A Synopsis of Aneura

Aneuraceae, Marchantiophyta — 54 accepted species

Aneura Dumort., Comment. Bot.: 115. 1822. (Dumortier 1822)
- =Trichostylium Corda, Deutschl. Fl., Abt. II, Cryptog. 26–27: 116. 1835. (Yano & Gradstein 1997)
- =Cryptothallus Malmb., Ann. Bryol. 6: 122. 1933. (Malmborg 1933; Crandall-Stotler et al. 2009)
Aneura amboinensis Steph., Bull. Herb. Boissier 7(9): 678. 1899. (Stephani 1899b)
Aneura augustae Steph., Sp. Hepat. (Stephani) 6: 430. 1923. (Stephani 1925)
Aneura biflora Colenso, Trans. & Proc. New Zealand Inst. 17: 262. 1885. (Colenso 1885)
Aneura blasioides (Horik.) Furuki, J. Hattori Bot. Lab. 70: 311. 1991. (Furuki 1991)

Basionym: Riccardia blasioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 197. 1933. (Horikawa 1933)
Aneura brasiliensis (Ångstr.) Steph., Bull. Herb. Boissier 7(10): 764. 1899. (Stephani 1899a)

Basionym: Pseudoneura brasiliensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33(7): 91. 1876. (Ångström 1876)
Aneura breutelii Steph., Bull. Herb. Boissier 7(10): 759. 1899. (Stephani 1899a; Rico Gutiérrez 2006)
Aneura brevifolia (Gottsche & Rabenh.) Steph., Hedwigia 32(1): 27. 1893. (Stephani 1893)

Basionym: Pseudoneura brevifolia Gottsche & Rabenh., Hepat. Eur.: 560. 1873. (Gottsche & Rabenhorst 1873)

≡Acrostolia brevifolia (Gottsche & Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 431. 1877. (Trevisan de Saint-Léon 1877)
Aneura brevissima Steph., Sp. Hepat. (Stephani) 6: 21. 1917. (Stephani 1925)
Aneura cerebrata Hewson, Proc. Linn. Soc. New South Wales 94: 185. 1970. (Hewson 1970)
Aneura crateriformis Furuki & D.G. Long, J. Bryol. 18: 281. 1994. (Furuki & Long 1994)
Aneura crinita C. Massal., Bull. Soc. Bot. Ital. 1917(8–9): 81. 1917. (Massalongo 1917)
Aneura crumii L. Söderstr., A. Hagborg & von Konrat, Phytotaxa 65: 43. 2012. (Söderström et al. 2012)

Replaced synonym: Cryptothallus hirsutus H.A. Crum, Bryologist 99: 438. 1996. (Crum & Bruce 1996)
Aneura densa Steph., Sp. Hepat. (Stephani) 6: 24. 1917. (Stephani 1925)
Aneura denticulata Mitt. ex Thurn, Timehri 5: 222. 1886. (Thurn 1886)
Aneura eachamensis Hewson, Proc. Linn. Soc. New South Wales 94: 184. 1970. (Hewson 1970)
Aneura erronea Steph., Sp. Hepat. (Stephani) 6: 20. 1917. (Stephani 1925)
Aneura eskuchei Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 294. 1986. (Hässel de Menéndez 1986)
Aneura gemmifera Furuki, J. Hattori Bot. Lab. 70: 309. 1991. (Furuki 1991)
Aneura giangena Hewson, Proc. Linn. Soc. New South Wales 94: 190. 1970. (Hewson 1970)
Aneura gibbsiana Steph., Sp. Hepat. (Stephani) 6: 28. 1917. (Stephani 1925)
Aneura glaucescens Steph., Sp. Hepat. (Stephani) 6: 28. 1917. (Stephani 1925)
Aneura goebeliana Steph., Sp. Hepat. (Stephani) 6: 28. 1917. (Stephani 1925)
Aneura hirsuta Furuki, J. Hattori Bot. Lab. 70: 317. 1991. (Furuki 1991)
Aneura hunsteinii Steph., Sp. Hepat. (Stephani) 6: 431. 1923. (Stephani 1925)
Aneura imbricata Colenso, Trans. & Proc. New Zealand Inst. 16: 359. 1884. (Colenso 1884)
Aneura intermedia Steph., Bull. Herb. Boissier 7(9): 692. 1899. (Stephani 1899b; Gradstein & Costa 2003)
Aneura kaguaensis Hewson, Proc. Linn. Soc. New South Wales 94: 190. 1970. (Hewson 1970)
Aneura keniae Gola, Mem. Reale Accad. Sci. Torino, ser. 2, 65: 3. 1914. (Gola 1914)
Aneura latemultifida Steph., Sp. Hepat. (Stephani) 6: 431. 1923. (Stephani 1925)
Aneura latissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544. 1885. (Spruce 1884–1885; Reeb & Gradstein 2020)

≡Riccardia latissima (Spruce) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Kl. 67: 177. 1898.
- =Riccardia pseudopinguis Herzog, Beih. Bot. Centralbl., Abt. 2, 61/B: 560. 1941[1942]. (Reeb & Gradstein 2020) ≡Aneura pseudopinguis (Herzog) Pócs, Acta Bot. Hung. 29: 133. 1983. (Gradstein et al. 1983; Reeb & Gradstein 2020)
Aneura ledermannii Steph., Sp. Hepat. (Stephani) 6: 431. 1923. (Stephani 1925)
Aneura macrostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550. 1885. (Spruce 1884–1885)
Aneura marianensis Furuki, Bryologist 97: 87. 1994. (Furuki 1994)
Aneura maxima (Schiffn.) Steph., Bull. Herb. Boissier 7: 668. 1899. (Stephani 1899b; Söderström et al. 2023)

Basionym: Riccardia maxima Schiffn., Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Kl. 67: 178. 1898. (Schiffner 1899)
Aneura mirabilis (Malmb.) Wickett & Goffinet, Bot. J. Linn. Soc. 156: 11. 2008. (Wickett & Goffinet 2008)

Basionym: Cryptothallus mirabilis Malmb., Ann. Bryol. 6: 122. 1933. (Malmborg 1933)
Aneura novaecaledoniae R.M. Schust., Phytologia 56: 451. 1985. (Schuster 1985)
Aneura novaguineensis Hewson, Proc. Linn. Soc. New South Wales 94: 189. 1970. (Hewson 1970)
- =Aneura gigantea R.M. Schust., Phytologia 56: 451. 1985, nom. illeg. (Schuster 1985; Brown & Braggins 1989)
Aneura nymanii Steph., Sp. Hepat. (Stephani) 6: 35. 1917. (Stephani 1925)
Aneura pellioides (Horik.) Inoue, J. Hattori Bot. Lab. 25: 209. 1962. (Inoue 1962; Söderström et al. 2023)

Basionym: Riccardia pellioides Horik., Bot. Mag. (Tokyo) 51: 429. 1937. (Horikawa 1937; Söderström et al. 2023)

≡Trichostylium pellioides (Horik.) R.M. Schust., Bryologist 61: 53. 1958.
Aneura pellucida Colenso, Trans. & Proc. New Zealand Inst. 18: 252. 1886. (Colenso 1886)
Aneura pinguis (L.) Dumort., Syll. Jungerm. Europ.: 86. 1831. (Dumortier 1831; Long et al. 2023)

Basionym: Jungermannia pinguis L., Sp. Pl. 2: 1136. 1753. (Linnaeus 1753; Reeb & Gradstein 2020)

≡Roemeria pinguis (L.) Raddi, Jungermanniogr. Etrusca: 37. 1818. ≡Riccardia pinguis (L.) Gray, Nat. Arr. Brit. Pl. 1: 684. 1821. (Paton 1999) ≡Blasia pinguis (L.) Fr., Stirp. Agri Femsion. 3: 31. 1825. (Fries 1825) ≡Metzgeria pinguis (L.) Corda, Naturalientausch 12 [Opiz, Beitr. Naturgesch.]: 654. 1829. ≡Gymnomitrion pingue (L.) Huebener, Hepaticol. Germ.: 41. 1834. (Huebener 1834; Schuster 1992) ≡Sarcomitrium pingue (L.) Mitt., Fl. Tasman. 2: 239. 1860[1859]. ≡Trichostylium pingue (L.) R.M. Schust., Bryologist 61: 53. 1958.
- =Jungermannia pinguis var. angustior Hook., Brit. Jungermann.: 46. 1813. ≡Riccardia pinguis var. angustior (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 684. 1821. ≡Aneura pinguis var. angustior (Hook.) Dumort., Syll. Jungerm. Europ.: 86. 1831. (Dumortier 1831; Hodgetts et al. 2020) ≡Aneura pinguis f. angustior (Hook.) Müll. Frib., Lebermoose 6(6): 332. 1908. ≡Riccardia pinguis f. angustior (Hook.) Nis Jensen, Mitt. Arbeitsgem. Florist. Schleswig-Holstein 4: 14. 1952.
- =Roemeria pinguis var. major Raddi, Jungermanniogr. Etrusca: 37 (48). 1818. (Schuster 1992)
- =Roemeria pinguis var. media Raddi, Jungermanniogr. Etrusca: 37 (48). 1818. (Schuster 1992)
- =Roemeria pinguis var. minor Raddi, Jungermanniogr. Etrusca: 37 (48). 1818. (Schuster 1992)
- =Jungermannia sessilis Spreng., Syst. Veg. [Sprengel] 4(1): 232. 1827. (Sprengel 1827) ≡Aneura sessilis (Spreng.) Gottsche, Lindenb. & Nees, Syn. Hepat. (fasc. 4): 495. 1846. (Gottsche et al. 1844; Schuster 1992) ≡Riccardia sessilis (Spreng.) Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 431. 1877.
- =Gymnomitrion pingue var. angustum Huebener, Hepaticol. Germ.: 41. 1834. (Huebener 1834; Schuster 1992)
- =Trichostylium affine Corda, Deutschl. Fl., Abt. II, Cryptog. 26–27: 116. 1835. (Schuster 1992)
- =Aneura pinguis f. crassior Nees, Naturgesch. Eur. Leberm. 3: 427. 1838. ≡Riccardia pinguis var. crassior (Nees) Dalla Torre & Sarnth., Fl. Tirol 5: 12. 1902. ≡Riccardia pinguis f. crassior (Nees) Schiffn., Krit. Bemerk. Eur. Leberm. 26: 12. 1941. (Schiffner 1941)
- =Aneura pinguis f. crispa Nees, Naturgesch. Eur. Leberm. 3: 428. 1838. ≡Riccardia pinguis f. crispa (Nees) Schiffn., Krit. Bemerk. Eur. Leberm. 26: 13. 1941. (Schiffner 1941)
- =Aneura pinguis f. denticulata Nees, Naturgesch. Eur. Leberm. 3: 428. 1838. ≡Aneura pinguis var. denticulata (Nees) Debat, Fl. Musc.: 235. 1874. (Paton 1999) ≡Riccardia pinguis var. denticulata (Nees) Besch. & C. Massal., Miss. Sci. Cape Horn, Bot. 5: 243. 1889.
- =Aneura pinguis f. prolifera Nees, Naturgesch. Eur. Leberm. 3: 428. 1838. (Nees von Esenbeck 1838)
- =Aneura pinguis f. breviseta Nees, Naturgesch. Eur. Leberm. 3: 429. 1838. (Nees von Esenbeck 1838)
- =Aneura pinguis f. fasciata Nees, Naturgesch. Eur. Leberm. 3: 429. 1838, nom. illeg. (Nees von Esenbeck 1838) ≡Riccardia pinguis var. fasciata (Nees) Schiffn., Sitzungsber. Deutsch. Naturwiss.-Med. Vereins Böhmen "Lotos" Prag 48: 322. 1900, nom. illeg.
- =Trichostylium arenarium Meyen, Arch. Anat. Physiol. Wiss. Med.: 273. 1839. (Schuster 1992)
- =Jungermannia rigida Wallr., Erst. Beitr. Fl. Hercyn.: 315. 1840, nom. illeg. (Wallroth 1840; Ros et al. 2007)
- =Aneura pinguis var. minor Gottsche, Lindenb. & Nees, Syn. Hepat. (Fasc. 4): 494. 1846. (Söderström et al. 2021)
- =Aneura pinguis var. exasperata Gottsche, Lindenb. & Nees, Syn. Hepat.: 494. 1846. (Gottsche et al. 1844)
- =Riccardia pinguis f. major Lindb., Hepaticol. Utveckl.: 37. 1877. (Isoviita 1970)
- =Riccardia pinguis f. minor Lindb., Hepaticol. Utveckl.: 37. 1877. (Isoviita 1970)
- =Riccardia fuscovirens Lindb., Helsingfors Dagblad 1878(84): 2. 1878. (Söderström et al. 2022) ≡Aneura fuscovirens (Lindb.) Steph., Hedwigia 32(3): 137. 1893. (Söderström et al. 2022) ≡Aneura pinguis f. fuscovirens (Lindb.) Kavina, Arch. Prir. Vyzk. Cech 16(2): 185. 1915. ≡Riccardia pinguis var. fuscovirens (Lindb.) C.E.O. Jensen, Danmarks Mosser 1: 63. 1915. ≡Riccardia pinguis subsp. fuscovirens (Lindb.) Schiffn., Krit. Bemerk. Eur. Leberm. 26: 15. 1941. ≡Aneura pinguis var. fuscovirens (Lindb.) Damsh., Ill. Fl. Nord. Liverw. Hornw.: 656. 2002. (Damsholt 2002)
- =Riccardia pinguis var. tenuis Nees ex Schiffn., Consp. Hepat. Arch. Ind.: 56. 1898. (Söderström et al. 2010) ≡Riccardia pinguis f. tenuis (Nees ex Schiffn.) Schiffn., Krit. Bemerk. Eur. Leberm. 26: 13. 1941.
- =Riccardia viridissima Schiffn., Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Kl. 67: 176. 1898. ≡Aneura viridissima (Schiffn.) Steph., Bull. Herb. Boissier 7(10): 763. 1899. (Thouvenot et al. 2011)
- =Aneura fuscovirens var. submersa Loeske ex Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfm.: 108. 1903[1902]. (Warnstorf 1902–1903)
- =Aneura pinguis var. glauca Kavina, Arch. Prir. Vyzk. Cech 16(2): 185. 1915. (Kavina 1915)
- =Aneura pinguis f. linearis Kavina, Arch. Prir. Vyzk. Cech 16(2): 185. 1915. (Kavina 1915)
- =Aneura pinguis f. calcarea Kavina, Arch. Prir. Vyzk. Cech 16(2): 186. 1915. (Kavina 1915)
- =Riccardia fuscovirens var. subcylindrica Lindb. ex Arnell & C.E.O. Jensen, Bot. Not. 1915: 190. 1915. (Arnell & Jensen 1915)
- =Aneura indica Steph., J. Bombay Nat. Hist. Soc. 25: 280. 1917. (Srivastava & Udar 1976)
- =Aneura laurentiana Steph., Sp. Hepat. (Stephani) 6: 32. 1917. (Wigginton 2018) ≡Riccardia laurentiana (Steph.) Demaret, Bull. Jard. Bot. État Bruxelles 16: 306. 1942.
- =Riccardia submarginata S.W. Arnell, Bot. Not. 1952: 139. 1952. (Arnell 1952; Wigginton 2018)
Aneura polyantha Colenso, Trans. & Proc. New Zealand Inst. 17: 262. 1885. (Colenso 1885)
Aneura puiggarii Steph., Sp. Hepat. (Stephani) 6: 39. 1917. (Stephani 1925; Gradstein & Costa 2003)
Aneura punctata Colenso, Trans. & Proc. New Zealand Inst. 18: 254. 1886. (Colenso 1886)
Aneura rodwayi Hewson, Proc. Linn. Soc. New South Wales 94: 188. 1970. (Hewson 1970)
Aneura roraimensis Steph., Trans. Linn. Soc. London, Bot. 6: 94. 1901. (Stephani 1901)
Aneura rotangicola Steph., Sp. Hepat. (Stephani) 6: 432. 1923. (Stephani 1925)
Aneura serrulata Gottsche ex Steph., Sp. Hepat. (Stephani) 6: 42. 1917. (Stephani 1925)
Aneura sharpii Inoue & N.G. Mill., Bull. Natl. Sci. Mus. Tokyo, B 11: 96. 1985. (Inoue & Miller 1985; Söderström et al. 2023)
Aneura singalangana (Schiffn.) Steph., Bull. Herb. Boissier 7(10): 751. 1899. (Stephani 1899a)

Basionym: Riccardia singalangana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Kl. 67: 174. 1898. (Schiffner 1899)
Aneura subcanaliculata R.M. Schust., J. Hattori Bot. Lab. 67: 60. 1989. (Schuster 1989)
Aneura subledermannii Steph., Sp. Hepat. (Stephani) 6: 432. 1923. (Stephani 1925)
Aneura subtenerrima Steph., Sp. Hepat. (Stephani) 6: 432. 1923. (Stephani 1925)
Aneura vincentina Steph., Sp. Hepat. (Stephani) 6: 45. 1917. (Stephani 1925)

A Synopsis of Aneura

References (68)