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A Classification of the Anthocerotophyta
28 accepted genera
Anthocerotophyta
Stotler & Crand.-Stotl.
, Bryologist 80(3): 425. 1977. (
Stotler & Crandall-Stotler 1977
)
Anthocerotopsida
Jancz.
, Bot. Zeitung (Berlin) 30(22): 413. 1872. (
Janczewski 1872
)
Anthocerotidae
Engl.
, Nat. Pflanzenfam. 1(3): 1, 6. 1893. (
Engler 1893
)
Anthocerotales
Rabenh.
, Krypt.-Fl. Sachsen: 299, 302. 1863. (
Rabenhorst 1863
)
Anthocerotaceae
Dumort.
, Anal. Fam. Pl.: 68. 1829. (
Dumortier 1829
)
=
Carpocerotaceae
Dumort. ex Nees
, Fl. Bras. Enum. Pl. 1(1): 303. 1833,
nom. illeg.
(
Martius et al. 1833
;
Grolle 1972
)
†
Anthocerisporis
Krutzsch
, Atlas Mittel- und Jungtert. Disp. Spor. 2: 3. 1963. (
Krutzsch 1963
;
Zhang et al. 2021
)
†
Anthoceroites
Khursel & Narkhede
, Int. J. Res. Biosci., Agric. Technol. 5(Special Issue, 2): 1198. 2017. (
Khursel & Narkhede 2017
)
Anthoceros
L.
, Sp. Pl. 2: 1139. 1753. (
Linnaeus 1753
)
=
Ceranthus
L. ex Gilib.
, Syst. Pl. Eur. 2(3): 466. 1787,
nom. illeg.
(
Gilibert 1787
;
Yano & Gradstein 1997
)
=
Carpoceros
Dumort.
, Comment. Bot.: 68, 76. 1822. (
Dumortier 1822
;
Yano & Gradstein 1997
)
=
Aspiromitus
Steph.
, Sp. Hepat. (Stephani) 5: 957. 1916. (
Stephani 1917
;
Grolle 1983
)
=
Folioceros
D.C. Bhardwaj
, Geophytology 1: 9. 1971. (
Bhardwaj 1971
;
Xu et al. 2025
)
=
Sphaerosporoceros
Hässel
, J. Hattori Bot. Lab. 64: 79. 1988. (
Hässel de Menéndez 1988
)
†
Bohemiasporis
Krutzsch
, Atlas Mittel- und Jungtert. Disp. Spor. 4–5: 16. 1967. (
Krutzsch 1967
;
Stuchlik et al. 2001
)
†
Rudolphisporis
Krutzsch
, Atlas Mittel- und Jungtert. Disp. Spor. 2: 4. 1963. (
Krutzsch 1963
;
Stuchlik et al. 2001
)
≡
†
Rudolphisporites
E. Nagy
, Acta Bot. Acad. Sci. Hung. 14: 114. 1968,
nom. illeg.
†
Saxosporis
Krutzsch
, Atlas Mittel- und Jungtert. Disp. Spor. 2: 4. 1963. (
Krutzsch 1963
;
Zhang et al. 2021
)
≡
†
Saxosporites
E. Nagy
, Acta Bot. Acad. Sci. Hung. 14: 113. 1968,
nom. illeg.
†
Sporollites
Pundkar
, Int. J. Adv. Innov. Res. 8(4): 27. 2021. (
Pundkar 2021
)
Dendrocerotidae
R.J. Duff, J.C. Villarreal, Cargill & Renzaglia
, Bryologist 110: 240. 2007. (
Duff et al. 2007
)
Dendrocerotales
T. Katag. & Furuki
, Hattoria 9: 81. 2018. (
Katagiri & Furuki 2018
)
Dendrocerotaceae
J. Haseg.
, J. Hattori Bot. Lab. 64: 93. 1988. (
Hasegawa 1988
)
Dendroceros
Nees
, Syn. Hepat.: 579. 1846. (
Gottsche et al. 1844
)
Megaceros
Campb.
, Ann. Bot. (Oxford) 21: 484. 1907. (
Campbell 1907
)
Nothoceros
(R.M. Schust.) J. Haseg.
, J. Hattori Bot. Lab. 76: 32. 1994 (‘
Notoceros
’). (
Hasegawa 1994
)
Basionym:
Megaceros
subg.
Nothoceros
R.M. Schust.
, Phytologia 63: 200. 1987. (
Schuster 1987
)
Phaeomegaceros
R.J. Duff, J.C. Villarreal, Cargill & Renzaglia
, Bryologist 110(2): 241. 2007. (
Duff et al. 2007
)
Phymatocerotales
R.J. Duff, J.C. Villarreal, Cargill & Renzaglia
, Bryologist 110(2): 240. 2007. (
Duff et al. 2007
)
Phymatocerotaceae
R.J. Duff, J.C. Villarreal, Cargill & Renzaglia
, Bryologist 110: 240. 2007. (
Duff et al. 2007
)
Phymatoceros
Stotler, W.T. Doyle & Crand.-Stotl.
, Phytologia 87(2): 114. 2005. (
Stotler et al. 2005
)
Notothyladidae
R.J. Duff, J.C. Villarreal, Cargill & Renzaglia
, Bryologist 110: 240. 2007. (
Duff et al. 2007
)
Notothyladales
Hyvönen & Piippo
, J. Hattori Bot. Lab. 74: 117. 1993. (
Hyvönen & Piippo 1993
)
Notothyladaceae
Müll. Frib. ex Prosk.
, Phytomorphology 10: 10. 1960. (
Proskauer 1960
)
=
Phaeocerotaceae
D.C. Bhardwaj
, Recent Advances Cryptog. Bot. 1: 149. 1982. (
Bhardwaj 1982
;
Hässel de Menéndez 1988
)
†
Annulispora
Jersey
, Queensland Gov. Mining J. 60: 358. 1959. (
Jersey 1959
;
Zhang et al. 2021
)
=
†
Distalanulisporites
Klaus
, Jahrb. Geol. Bundesanst. Sonderb. 5: 131. 1960. (
Klaus 1960
;
Kavary 1972
)
=
†
Bianulisporites
Pautsch
, Acta Palaeobot. 12: 19. 1971. (
Pautsch 1971
;
McKellar 1974
)
†
Asterisporites
Venkatach. & Rawat
, Palaeobotanist 19(1–3): 106. 1970[1971]. (
Venkatachala & Rawat 1970
;
Zhang et al. 2021
)
=
†
Trisolissporites
B.D. Tschudy
, Geol. Surv. Profess. Pap. 770: 8. 1973,
nom. illeg.
(
Tschudy 1973
;
Li 2001
)
†
Capsulites
Saksena
, J. Indian Bot. Soc. 26: 246. 1948. (
Saksena 1948
;
Oostendorp 1987
)
†
Dicyclosporis
(Eb. Schulz) Janson. & Hills
, Genera File Foss. Spor. Pollen (Suppl. 10): 4642. 1990. (
Jansonius & Hills 1990
;
Zhang et al. 2021
)
Basionym:
†
Stereisporites
subg.
Dicyclosporis
Eb. Schulz
, Beih. Z. Geol. 55: 78. 1966. (
Döring et al. 1966
)
†
Distcyclosporis
(Eb. Schulz) McKellar
, Geol. Surv. Queensland Publ. 361: 31. 1974. (
McKellar 1974
;
Zhang et al. 2021
)
Basionym:
†
Stereisporites
subg.
Distcyclosporis
Eb. Schulz
, Beih. Z. Geol. 55: 79. 1966. (
Döring et al. 1966
)
†
Foraminisporis
Krutzsch
, Beih. Z. Geol. 21–22: 130. 1959. (
Krutzsch 1959
;
Zhang et al. 2021
)
Mesoceros
Piippo
, Acta Bot. Fenn. 148: 30. 1993. (
Piippo 1993
)
†
Nevesisporites
Jersey & Paten
, Geol. Surv. Queensland Publ. 322: 8. 1964. (
Jersey & Paten 1964
;
Zhang et al. 2021
)
†
Notothylacites
Němejc & Pacltová
, Palaeobotanist 21: 24. 1974. (
Němejc & Pacltová 1974
;
Oostendorp 1987
)
Notothylas
Sull.
, Amer. J. Sci. Arts, ser. 2, 1: 74. 1846. (
Gray 1846
)
Replaced synonym:
Carpobolus
Schwein.
, J. Acad. Nat. Sci. Philadelphia 2: 366. 1822,
nom. illeg.
(
Schweinitz 1822
;
Yano & Gradstein 1997
)
≡
Carpolipum
Nees
, Syn. Hepat.: 591. 1846,
nom. illeg.
(
Yano & Gradstein 1997
)
=
Chamaeceros
Milde
, Bot. Zeitung (Berlin) 14: 767. 1856. (
Milde 1856
;
Yano & Gradstein 1997
)
Paraphymatoceros
Hässel
, Phytologia 88: 208. 2006. (
Hässel de Menéndez 2006
)
Phaeoceros
Prosk.
, Bull. Torrey Bot. Club 78(4): 346. 1951. (
Proskauer 1951
)
=
Phaeoceros
subg.
Hattorioceros
J. Haseg.
, J. Hattori Bot. Lab. 75: 272. 1992. (
Hasegawa 1994
)
≡
Hattorioceros
(J. Haseg.) J. Haseg.
, J. Hattori Bot. Lab. 76: 32. 1994. (
Hasegawa 1994
;
Li et al. 2011
)
†
Phaeocerosporites
E. Nagy
, Acta Bot. Acad. Sci. Hung. 14(1–2): 116. 1968. (
Nagy 1968
;
Zhang et al. 2021
)
†
Simeonospora
Balme
, Univ. Kansas Paleontol. Contr., Special Publ. 4: 329. 1970. (
Balme 1970
;
Zhang et al. 2021
)
Leiosporocerotopsida
Stotler & Crand.-Stotl.
, Bryologist 108(1): 24. 2005. (
Stotler & Crandall-Stotler 2005
)
Basionym:
Leiosporocerotales
Hässel
, J. Hattori Bot. Lab. 64: 82. 1988. (
Hässel de Menéndez 1988
)
Leiosporocerotales
Hässel
, J. Hattori Bot. Lab. 64: 82. 1988. (
Hässel de Menéndez 1988
)
Leiosporocerotaceae
Hässel ex Ochyra
, Polish Bot. J. 56(1): 115. 2011. (
Ochyra 2011
)
†
Casterlorum
Retallack
, Palaeobotanist 68(1–2): 183. 2019. (
Retallack 2019
)
Leiosporoceros
Hässel
, J. Bryol. 14: 255. 1986. (
Hässel de Menéndez 1986
)
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