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A Synopsis of
Diplophyllum
Scapaniaceae, Marchantiophyta — 24 accepted species
Diplophyllum
(Dumort.) Dumort.
, Recueil Observ. Jungerm.: 15. 1835,
nom. cons.
(
Dumortier 1835
)
Basionym:
Jungermannia
sect.
Diplophyllum
Dumort.
, Syll. Jungerm. Europ.: 44. 1831. (
Dumortier 1831
)
≡
Diplophylleia
(Rchb.) Trevis.
, Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 420. 1877. (
Trevisan de Saint-Léon 1877
;
Yano & Gradstein 1997
)
=
Krunodiplophyllum
Grolle
, J. Hattori Bot. Lab. 28: 70. 1965. (
Engel & Merrill 1998
)
Diplophyllum africanum
S.W. Arnell
, Ark. Bot., n.s. 3(16): 527. 1956. (
Arnell 1956
)
Diplophyllum albicans
(L.) Dumort.
, Recueil Observ. Jungerm.: 16. 1835. (
Dumortier 1835
)
G5
Basionym:
Jungermannia albicans
L.
, Sp. Pl. 2: 1133. 1753. (
Linnaeus 1753
;
Damsholt 2002
)
≡
Scapania albicans
(L.) Rabenh.
, Krypt.-Fl. Sachsen 1: 338. 1863. (
Hodgetts et al. 2020
)
≡
Diplophylleia albicans
(L.) Trevis.
, Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4(13): 420. 1877. (
Trevisan de Saint-Léon 1877
;
Damsholt 2002
)
=
Jungermannia varia
L.
, Sp. Pl.: 1135. 1753. (
Linnaeus 1753
;
Schumacker & Váňa 2005
)
=
Jungermannia albicans
var.
procumbens
Hook.
, Brit. Jungermann.: pl. 25. 1812.
≡
Jungermannia albicans
f.
procumbens
(Hook.) Nees
, Naturgesch. Eur. Leberm. 1: 228. 1833.
≡
Diplophyllum albicans
var.
procumbens
(Hook.) Dumort.
, Bull. Soc. Roy. Bot. Belgique 13: 48. 1874.
≡
Diplophyllum albicans
f.
procumbens
(Hook.) Müll. Frib.
, Lebermoose 6(20): 357. 1914. (
Müller 1912-1916
)
=
Jungermannia falcata
Raddi
, Jungermanniogr. Etrusca: 22. 1818,
nom. illeg.
(
Ros et al. 2007
)
≡
Jungermannia albicans
var.
falcata
De Not.
, Mem. Reale Accad. Sci. Torino, ser. 2, 18: 463. 1859.
=
Jungermannia albicans
f.
infuscata
Nees
, Naturgesch. Eur. Leberm. 1: 228. 1833.
≡
Jungermannia albicans
var.
infuscata
(Nees) Gottsche, Lindenb. & Nees
, Syn. Hepat.: 76. 1844.
≡
Diplophyllum albicans
f.
infuscatum
(Nees) Schiffn.
, Krit. Bemerk. Eur. Leberm. 14: 17. 1919. (
Schiffner 1919
)
=
Jungermannia albicans
var.
conferta
Huebener
, Hepaticol. Germ.: 256. 1834. (
Huebener 1834
)
=
Jungermannia albicans
var.
rupestris
Huebener
, Hepaticol. Germ.: 256. 1834. (
Huebener 1834
)
=
Jungermannia albicans
var.
uliginosa
Huebener
, Hepaticol. Germ.: 256. 1834. (
Huebener 1834
)
=
Jungermannia fissidentoidea
Huebener
, Hepaticol. Germ.: 258. 1834. (
Ros et al. 2007
)
≡
Diplophyllum albicans
var.
fissidentoidea
(Huebener) Saut.
, Oesterr. Bot. Z. 44: 129. 1894.
=
Jungermannia albicans
var.
balsamita
Grognot
, Pl. Crypt. Saône-et-Loire: 42. 1863. (
Grognot 1863
)
=
Jungermannia albicans
var.
nervosa
Grognot
, Pl. Crypt. Saône-et-Loire: 42. 1863. (
Grognot 1863
)
=
Jungermannia albicans
var.
rubescens
Grognot
, Pl. Crypt. Saône-et-Loire: 42. 1863. (
Ros et al. 2007
)
=
Jungermannia albicans
var.
virens
Grognot
, Pl. Crypt. Saône-et-Loire: 42. 1863. (
Grognot 1863
)
=
Diplophyllum albicans
f.
propaguliferum
Corb.
, Mém. Soc. Sci. Nat. Math. Cherbourg 26: 330. 1889. (
Corbière 1889
)
≡
Jungermannia albicans
f.
propagulifera
(Corb.) Hérib.
, Mém. Acad. Sci. Clermont-Ferrand, sér. 2, 14: 470. 1899.
=
Jungermannia albicans
f.
alpina
Steph. ex Hérib.
, Mém. Acad. Sci. Clermont-Ferrand, sér. 2, 14: 470. 1899. (
Héribaud 1899
)
=
Diplophyllum albicans
var.
purpurascens
C.E.O. Jensen
, Bot. Faeröes 1: 130. 1901. (
Jensen 1901
;
Müller 1912-1916
)
≡
Diplophyllum albicans
f.
purpurascens
(C.E.O. Jensen) Müll. Frib.
, Lebermoose 6(20): 357. 1914. (
Müller 1912-1916
)
=
Diplophyllum albicans
var.
subacutum
Velen.
, Rozpr. České Akad. Císaře Františka Josefa Vědy, Tř. 2, Vědy Math. Přír. 10(12): 13. 1901. (
Velenovský 1901
)
=
Diplophyllum albicans
var.
fuscum
Loeske
, Moosfl. Harz.: 63. 1903. (
Loeske 1903
)
≡
Diplophyllum albicans
f.
fuscum
(Loeske) Müll. Frib.
, Lebermoose 6(20): 357. 1914.
=
Diplophyllum albicans
f.
aquaticum
Müll. Frib.
, Lebermoose 6(20): 357. 1914. (
Müller 1912-1916
)
=
Diplophylleia albicans
var.
elegans
C.E.O. Jensen
, Danmarks Mosser 1: 147. 1915. (
Jensen 1915
)
=
Diplophylleia albicans
var.
recurva
C.E.O. Jensen
, Danmarks Mosser 1: 147. 1915. (
Jensen 1915
)
=
Diplophylleia albicans
var.
secunda
C.E.O. Jensen
, Danmarks Mosser 1: 147. 1915. (
Jensen 1915
)
=
Diplophyllum albicans
var.
lucifugum
E. Bauer ex Schiffn.
, Krit. Bemerk. Eur. Leberm. 14: 18. 1919. (
Schiffner 1919
)
=
Diplophyllum albicans
f.
umbrosum
Schiffn.
, Krit. Bemerk. Eur. Leberm. 14: 18. 1919. (
Schiffner 1919
)
Diplophyllum andicola
R.M. Schust.
, Phytologia 39: 248. 1978. (
Schuster 1978
)
Diplophyllum andrewsii
A. Evans
, Bryologist 25(2): 28. 1922. (
Evans 1922
)
G3
Diplophyllum androgynum
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 277. 1998. (
Engel & Merrill 1998
)
Diplophyllum angustifolium
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 262. 1998. (
Engel & Merrill 1998
)
Diplophyllum apiculatum
(A. Evans) Steph.
, Sp. Hepat. (Stephani) 4: 110. 1910. (
Stephani 1912
)
G5
Basionym:
Diplophylleia apiculata
A. Evans
, Bot. Gaz. 34(5): 372. 1902. (
Evans 1902
)
Diplophyllum dioicum
R.M. Schust.
, Bull. Natl. Sci. Mus. Tokyo, n.s. 11: 20. 1968. (
Schuster 1968
)
Diplophyllum exiguum
Steph.
, Sp. Hepat. (Stephani) 6: 500. 1924. (
Stephani 1925
)
Diplophyllum gemmiparum
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 255. 1998. (
Engel & Merrill 1998
)
Diplophyllum incrassatum
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 265. 1998. (
Engel & Merrill 1998
)
Diplophyllum kinabaluense
Furuki & Suleiman
, J. Jap. Bot. 91(Suppl.): 340. 2016. (
Furuki & Suleiman 2016
)
Diplophyllum nanum
Herzog
, Memoranda Soc. Fauna Fl. Fenn. 26: 48. 1951. (
Herzog 1951
)
Diplophyllum novum
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 274. 1998. (
Engel & Merrill 1998
)
Diplophyllum obtusatum
(R.M. Schust.) R.M. Schust.
, Hepat. Anthocerotae N. Amer. 3: 215. 1974. (
Schuster 1974
)
G2
Basionym:
Diplophyllum apiculatum
var.
obtusatum
R.M. Schust.
, Amer. Midl. Naturalist 49(2): 432. 1953. (
Schuster 1953
)
Diplophyllum obtusifolium
(Hook.) Dumort.
, Recueil Observ. Jungerm.: 16. 1835. (
Dumortier 1835
)
G4
Basionym:
Jungermannia obtusifolia
Hook.
, Brit. Jungermann.: pl. 26. 1816[1812]. (
Hooker 1816
;
Damsholt 2002
)
≡
Scapania obtusifolia
(Hook.) Mitt.
, Fl. Tasman. 2: 233. 1859.
≡
Diplophylleia obtusifolia
(Hook.) Trevis.
, Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4(13): 420. 1877. (
Trevisan de Saint-Léon 1877
)
=
Jungermannia obtusifolia
var.
purpurascens
Lindenb.
, Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 14(Suppl.): 60. 1829.
≡
Diplophyllum obtusifolium
var.
purpurascens
(Lindenb.) Dalla Torre & Sarnth.
, Fl. Tirol 5: 75. 1902. (
Dalla Torre & Sarnthein 1903
)
=
Jungermannia obtusifolia
f.
exigua
Nees
, Naturgesch. Eur. Leberm. 1: 237. 1833. (
Nees von Esenbeck 1833
)
≡
Jungermannia obtusifolia
var.
exigua
(Nees) Debat
, Fl. Musc.: 216. 1874.
≡
Diplophyllum obtusifolium
f.
exiguum
(Nees) Dalla Torre & Sarnth.
, Fl. Tirol 5: 75. 1904.
=
Jungermannia obtusifolia
f.
tenera
Nees
, Naturgesch. Eur. Leberm. 1: 237. 1833. (
Nees von Esenbeck 1833
)
≡
Jungermannia obtusifolia
var.
tenera
(Nees) Debat
, Fl. Musc.: 216. 1874.
=
Jungermannia obtusifolia
var.
denticulata
Huebener
, Hepaticol. Germ.: 261. 1834,
nom. illeg.
≡
Diplophyllum obtusifolium
var.
denticulatum
(Huebener) Dumort.
, Bull. Soc. Roy. Bot. Belgique 13: 51. 1874,
nom. illeg.
(
Dumortier 1874
)
=
Jungermannia obtusifolia
var.
denticulata
Hartm.
, Handb. Skand. Fl. (ed. 3) 2: 324. 1838,
nom. illeg.
(
Hartman 1838
)
=
Jungermannia obtusifolia
var.
americana
Gottsche
, Linnaea 28(5): 549. 1856[1857]. (
Gottsche 1856
)
=
Jungermannia obtusifolia
var.
corsicana
Gottsche
, Linnaea 28(5): 549. 1856[1857]. (
Gottsche 1856
)
=
Scapania microscopica
Culm.
, Bull. Soc. Bot. France 62: 54. 1915. (
Ros et al. 2007
)
Diplophyllum obtusifolium
subsp.
domesticum
(Gottsche) Váňa
, Phytotaxa 76(3): 29. 2013. (
Váňa et al. 2013
)
Basionym:
Jungermannia domestica
Gottsche
, Linnaea 28: 548. 1856[1857]. (
Gottsche 1856
)
≡
Diplophyllum domesticum
(Gottsche) Steph.
, Hedwigia 33: 6. 1894. (
Váňa et al. 2013
)
=
Jungermannia obtusifolia
var.
procerior
Gottsche
, Frag. Suppl.: 53. 1880. (
Schiffner 1893
)
≡
Diplophyllum obtusifolium
var.
procerius
(Gottsche) Schiffn.
, Hepat. (Engl.-Prantl): 112. 1893.
=
Diplophyllum acutilobum
Steph.
, Kongl. Svenska Vetensk. Acad. Handl., n.s. 46(9): 83. 1911. (
Váňa et al. 2013
)
=
Diplophyllum marionense
S.W. Arnell
, Svensk Bot. Tidskr. 47(3): 415. 1953. (
Arnell 1953
;
Váňa et al. 2013
)
=
Diplophyllum domesticum
var.
icari
J.J. Engel & G.L. Merr.
, J. Hattori Bot. Lab. 84: 273. 1998. (
Engel & Merrill 1998
;
Engel & Glenny 2008
)
Diplophyllum obtusifolium
(Hook.) Dumort. subsp.
obtusifolium
Diplophyllum purpurascens
Bakalin & Vilnet
, Phytotaxa 447(2): 120. 2020. (
Bakalin et al. 2020
)
Diplophyllum recurvifolium
C. Massal.
, Atti Reale Ist. Veneto Sci. Lett. Arti 87: 221. 1928. (
Massalongo 1928
)
Diplophyllum serrulatum
(Müll. Frib.) Steph.
, Sp. Hepat. (Stephani) 4: 112. 1910. (
Stephani 1912
)
Basionym:
Diplophylleia serrulata
Müll. Frib.
, Bull. Herb. Boissier, sér. 2, 3(1): 34. 1903[1902]. (
Müller 1903
)
=
Diplophyllum sendaicum
Steph.
, Sp. Hepat. (Stephani) 6: 500. 1924. (
Yamada & Iwatsuki 2006
)
Diplophyllum sibiricum
Vilnet & Bakalin
, Pl. Syst. Evol. 304(10): 1280. 2018. (
Bakalin & Vilnet 2018
)
Diplophyllum squarrosum
Steph.
, Sp. Hepat. (Stephani) 4: 116. 1910. (
Stephani 1912
)
≡
Krunodiplophyllum squarrosum
(Steph.) Grolle
, J. Hattori Bot. Lab. 28: 71. 1965.
≡
Blepharidophyllum squarrosum
(Steph.) R.M. Schust.
, Bull. Natl. Sci. Mus. Tokyo, n.s. 14(4): 655. 1971.
Diplophyllum taxifolium
(Wahlenb.) Dumort.
, Recueil Observ. Jungerm.: 16. 1835. (
Dumortier 1835
)
G5
Basionym:
Jungermannia taxifolia
Wahlenb.
, Fl. Lapp.: 389. 1812. (
Wahlenberg 1812
)
≡
Jungermannia albicans
var.
taxifolia
(Wahlenb.) Wallr.
, Fl. Crypt. Germ. 1: 58. 1831. (
Wallroth 1831
)
≡
Jungermannia albicans
f.
taxifolia
(Wahlenb.) Nees
, Naturgesch. Eur. Leberm. 1: 228. 1833.
≡
Jungermannia albicans
subsp.
taxifolia
(Wahlenb.) Ångstr.
, Disp. Musc. Scand.: 25. 1842.
≡
Scapania albicans
var.
taxifolia
(Wahlenb.) Rabenh.
, Krypt.-Fl. Sachsen 1: 338. 1863.
≡
Diplophyllum albicans
var.
taxifolium
(Wahlenb.) Lindb.
, Contr. Fl. Crypt. As.: 236. 1872[1873].
≡
Diplophylleia taxifolia
(Wahlenb.) Trevis.
, Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4(13): 420. 1877. (
Trevisan de Saint-Léon 1877
)
≡
Diplophylleia albicans
var.
taxifolia
(Wahlenb.) C. Massal. & Carestia
, Nuovo Giorn. Bot. Ital. 12(4): 323. 1880.
=
Diplophyllum taxifolium
var.
macrostictum
H. Buch
, Commentat. Biol. 3(1): 23. 1928. (
Hodgetts et al. 2020
)
Diplophyllum taxifolium
var.
mucronatum
R.M. Schust.
, Hepat. Anthocerotae N. Amer. 3: 203. 1974. (
Schuster 1974
;
Stotler & Crandall-Stotler 2017
)
T1
Diplophyllum taxifolium
(Wahlenb.) Dumort. var.
taxifolium
Diplophyllum trollii
Grolle
, Khumbu Himal 1(4): 273. 1966. (
Grolle 1966
)
Diplophyllum verrucosum
R.M. Schust.
, Bull. Natl. Sci. Mus. Tokyo, n.s. 11: 19. 1968. (
Schuster 1968
)
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