A Synopsis of Radula

Radulaceae, Marchantiophyta — 247 accepted species

Radula Dumort., Comment. Bot.: 112. 1822, nom. cons. (Dumortier 1822)
- =Candollea Raddi, Jungermanniogr. Etrusca: 11. 1818, nom. illeg. (Yano & Gradstein 1997) ≡Patarola Léman, Dict. Sci. Nat. (ed. 2) 38: 73. 1825. (Yano & Gradstein 1997)
- =Martinellius Gray, Nat. Arr. Brit. Pl. 1: 690. 1821, nom. rej. (Yano & Gradstein 1997)
- =Patarola Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 434. 1877, nom. illeg. (Grolle 1983)
- =Stephanina Kuntze, Revis. Gen. Pl.: 839. 1891, nom. illeg. (Yano & Gradstein 1997)
Radula acuminata Steph., Sp. Hepat. (Stephani) 4: 230. 1910. (Stephani 1912)
- =Radula acuminata f. corticola S. Hatt., Bull. Tokyo Sci. Mus. 11: 81. 1944. (Yamada & Iwatsuki 2006)
- =Radula yunnanensis P.C. Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 39. 1955. (Zhu & So 2001)
Radula acuta Mitt., Fl. Vit.: 410. 1871[1873]. (Mitten 1873; So 2006)

≡Radula anceps var. subintegra Steph., Hedwigia 23(10): 156. 1884.
- =Radula mucronata Steph., Hedwigia 23: 114. 1884. (So 2006)
- =Radula apiculata Sande Lac. ex Steph., Hedwigia 23: 150. 1884. (So 2006) ≡Stephanina apiculata (Sande Lac. ex Steph.) Yasuda, Shokub. Kak.: 714. 1911.
- =Radula paucidens Steph. ex Castle, Rev. Bryol. Lichénol. 30: 39. 1961. (Söderström et al. 2012)
Radula acutangula Steph., Bull. Herb. Boissier 5: 848. 1897. (Stephani 1897b)
- =Radula pauciramea Steph., Sp. Hepat. (Stephani) 6: 513. 1924. (Staples & Imada 2006)
Radula acutiloba Steph., Hedwigia 28: 271. 1889. (Stephani 1889)
- =Radula papulosa Steph., J. Linn. Soc., Bot. 29: 272. 1892. (Renner 2014)
Radula aguirrei R.M. Schust. ex M.A.M. Renner, Phytotaxa 202(1): 70. 2015. (Söderström et al. 2015b)
Radula allisonii Castle, Rev. Bryol. Lichénol. 31(3–4): 148. 1962[1963]. (Castle 1962; Söderström et al. 2016)
Radula amentulosa Mitt., Bonplandia (Hannover) 9: 367. 1861. (Mitten 1861a)
- =Radula koghiensis K. Yamada, J. Hattori Bot. Lab. 58: 118. 1985. (Yamada 1985b; Thouvenot et al. 2011)
- =Radula recurvilobula K. Yamada, J. Hattori Bot. Lab. 58: 122. 1985. (Yamada 1985b; Thouvenot et al. 2011)
Radula amoena Herzog, Mitt. Inst. Allg. Bot. Hamburg 7: 192. 1931. (Herzog 1931)
Radula anceps Sande Lac., Ned. Kruidk. Arch. 3(4): 419. 1855. (Sande Lacoste 1855)
Radula aneurismalis (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 262. 1845. (Gottsche et al. 1844; So 2005a)

Basionym: Jungermannia aneurismalis Hook. f. & Taylor, London J. Bot. 4: 86. 1845. (Hooker & Taylor 1845)

≡Stephanina aneurismalis (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula compacta Castle, Rev. Bryol. Lichénol. 32: 43. 1963. (So 2005a)
Radula angulata Steph., Hedwigia 23: 114. 1884. (Stephani 1884b)
- =Radula bogotensis Steph., Hedwigia 23: 115. 1884. (Stephani 1884b; Gradstein 2021)
- =Radula korthalsii Steph., Hedwigia 23: 133. 1884. (Stephani 1884c; Gradstein & Uribe-M. 2016)
Radula anisotoma M.A.M. Renner, PhytoKeys 27(Special issue): 30. 2013. (Renner et al. 2013c)
Radula appressa Mitt., Philos. Trans. 168(extra vol.): 397. 1879. (Mitten 1879)
- =Radula silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349. 1882. (Gottsche 1882; Gradstein et al. 2022)
- =Radula guineensis Steph., Hedwigia 23(9): 133. 1884. (Wigginton 2018)
- =Radula comorensis Steph., Hedwigia 23: 132. 1884. (Stephani 1884c; Gradstein et al. 2022)
- =Radula mascarena Steph., Hedwigia 23: 134. 1884. (Wigginton 2018)
- =Radula oblongiloba Steph., Hedwigia 23: 153. 1884. (Wigginton 2018)
- =Radula angustata Steph., Bot. Jahrb. Syst. 8: 85. 1887[1886]. (Wigginton 2018)
- =Radula molleri Steph., Sp. Hepat. (Stephani) 4: 169. 1910. (Wigginton 2018)
Radula aquilegia (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 260. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia aquilegia Hook. f. & Taylor, London J. Bot. 3: 391 [as 291]. 1844. (Hooker & Taylor 1844a)

≡Martinellius aquilegius (Hook. f. & Taylor) Carruth., J. Bot. 3: 301. 1865. ≡Stephanina aquilegia (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Jungermannia complanata var. minor Hook., Brit. Jungermann.: tab. 81. 1816. (Castle 1968a) ≡Martinellius complanatus var. minor (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 691. 1821. ≡Radula complanata var. minor (Hook.) Dumort., Syll. Jungerm. Europ.: 38. 1831.
Radula assamica Steph., Hedwigia 23: 151. 1884. (Stephani 1884a)
- =Radula platyglossa P.C. Chen, Acta Phytotax. Sin. 9: 221. 1964. (Chen & Wu 1964; Zhu & So 2001)
Radula australiana K. Yamada, J. Hattori Bot. Lab. 51: 323. 1982. (Yamada 1982a)
Radula australis Austin, Bot. Bull. (Hanover) 1(7): 32. 1876. (Austin 1876; Stotler & Crandall-Stotler 2017)
- =Radula caloosiensis Austin, Bull. Torrey Bot. Club 6: 301. 1879. (Austin 1879; Schuster 1980)
- =Radula langloisii Castle, Bull. Torrey Bot. Club 52: 435. 1925. (Schuster 1980)
Radula bahiensis F.R. Oliveira-da-Silva, Ilk.-Borg. & Gradst., Phytotaxa 454(1): 25. 2020. (Oliveira-da-Silva et al. 2020)
†Radula baltica Heinrichs, Schäf.-Verw. & M.A.M. Renner, Rev. Palaeobot. Palynol. 235: 158. 2016. (Heinrichs et al. 2016)
Radula bischlerae Gradst. & Ilk.-Borg., Mem. New York Bot. Gard. 121: 623. 2021. (Gradstein 2021)
Radula bolanderi Gottsche ex Steph., Hedwigia 23(10): 145. 1884. (Stephani 1884a)
- =Radula spicata Austin, Bull. Torrey Bot. Club 6(3): 19. 1875, nom. illeg. (Austin 1875; Doyle & Stotler 2006)
- =Radula arctica Steph., Bot. Jahrb. Syst. 8: 98. 1886. (Söderström et al. 2015a)
Radula boninensis Furuki & K. Yamada, J. Jap. Bot. 61: 312. 1986. (Furuki & Yamada 1986)
Radula borneensis Steph., Sp. Hepat. (Stephani) 4: 209. 1910. (Stephani 1912)
Radula brasilica K. Yamada, J. Hattori Bot. Lab. 74: 35. 1993. (Yamada 1993b)
Radula buccinifera (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 261. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia buccinifera Hook. f. & Taylor, London J. Bot. 3: 580. 1844. (Hooker & Taylor 1844c)

≡Stephanina buccinifera (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891. ≡Patarola buccinifera (Hook. f. & Taylor) Trevis. ex Grolle, Acta Bot. Fenn. 121: 30. 1983.
- =Radula wattsiana Steph., Sp. Hepat. (Stephani) 4: 210. 1910. (Renner et al. 2013c)
Radula caduca K. Yamada, J. Hattori Bot. Lab. 45: 225. 1979. (Yamada 1979a)
Radula camerunensis Pócs & Döbbeler, Stud. Bot. Hung. 48(1): 24. 2017. (Pócs 2017; Wigginton 2018)
Radula campanulata Lindenb. & Gottsche, Syn. Hepat.: 256. 1844. (Gottsche et al. 1844)

≡Stephanina campanulata (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula pocillifera Taylor, London J. Bot. 5: 377. 1846. (Taylor 1846; Yamada 1993a) ≡Stephanina pocillifera (Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
Radula carringtonii J.B. Jack, Flora 64: 385. 1881. (Jack 1881; Hodgetts et al. 2020)

Replaced synonym: Radula aquilegia var. major Carrington, Trans. Bot. Soc. Edinburgh 7(3): 455. 1863. (Carrington 1863; Yamada 1986)
Radula castlei Grolle, Bryologist 73: 662. 1970. (Grolle 1970)
Radula cavifolia Hampe ex Gottsche, Lindenb. & Nees, Syn. Hepat.: 259. 1845. (Gottsche et al. 1844; Yamada 1979a)

≡Stephanina cavifolia (Hampe ex Gottsche, Lindenb. & Nees) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula magnilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 127. 1932. (Horikawa 1932; Yamada & Iwatsuki 2006)
Radula ceylanica K. Yamada, J. Jap. Bot. 50: 373. 1975. (Yamada 1975b)
Radula cochabambaensis K. Yamada, J. Hattori Bot. Lab. 74: 37. 1993. (Yamada 1993b)
Radula complanata (L.) Dumort., Syll. Jungerm. Europ.: 38. 1831. (Dumortier 1831)

Basionym: Jungermannia complanata L., Sp. Pl. 2: 1133. 1753. (Linnaeus 1753; Damsholt 2002)

≡Candollea complanata (L.) Raddi, Jungermanniogr. Etrusca: 13 (24). 1818. ≡Martinellius complanatus (L.) Gray, Nat. Arr. Brit. Pl. 1: 691. 1821. ≡Jubula complanata (L.) Corda, Deutschl. Fl., Abt. II, Cryptog. 26–27: 152. 1835. ≡Stephanina complanata (L.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Jungermannia epiphlaea Schrank, Naturhist. Briefe Oestreich 2: 305. 1785. (Schrank 1792)
- =Jungermannia cinerea Dicks., Fasc. Pl. Crypt. Brit. 2: 15. 1790. (Dickson 1790; Yamada 1979a)
- =Candollea complanata var. minor Raddi, Jungermanniogr. Etrusca: 14 (25). 1818. ≡Radula complanata var. minor (Raddi) Lindb., Hepaticol. Utveckl.: 35. 1877, nom. illeg. (Isoviita 1970)
- =Jungermannia complanata var. adscendens Hartm., Handb. Skand. Fl.: 438. 1820. (Hartman 1820)
- =Jungermannia complanata var. repens Hartm., Handb. Skand. Fl.: 438. 1820. (Hartman 1820)
- =Jungermannia complanata var. compacta Grognot, Pl. Crypt. Saône-et-Loire: 42. 1863. (Roumeguère 1888) ≡Radula complanata f. compacta (Grognot) Roum., Mém. Soc. Arts Sci. Carcassonne 5: 176. 1888.
- =Radula alpestris Lindb. ex Berggr., Bidr. Skand. Bryol.: 29. 1866. (Castle 1968a) ≡Radula complanata var. alpestris (Lindb. ex Berggr.) Lindb., Musci Scand.: 3. 1879. ≡Stephanina alpestris (Lindb. ex Berggr.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula hallii Austin, Bull. Torrey Bot. Club 6(3): 19. 1875. (Austin 1875; Schuster 1980)
- =Radula notarisii Steph., Hedwigia 23: 129. 1884. (Ros et al. 2007) ≡Radula complanata var. notarisii (Steph.) Schiffn., Hedwigia 48(3): 201. 1909.
- =Radula mauritiana Mitt. ex Steph., Hedwigia 23: 148. 1884. (Wigginton 2018)
- =Radula krausei Steph., Bot. Jahrb. Syst. 8: 97. 1886. (Söderström et al. 2015a)
- =Radula hyalina Steph., Sp. Hepat. (Stephani) 6: 511. 1924. (Yamada & Iwatsuki 2006)
Radula conferta Lindenb. & Gottsche, Syn. Hepat.: 729. 1847. (Gottsche et al. 1844)

≡Stephanina conferta (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula conferta var. laxior Gottsche, Mexik. Leverm.: 150 (246). 1863. (Gottsche 1863)
Radula constricta Steph., Sp. Hepat. (Stephani) 6: 506. 1924. (Stephani 1925; Bakalin & Klimova 2020)
Radula cordata Mitt., Fl. Vit.: 410. 1871[1873]. (Mitten 1873)
- =Radula mannii Austin, Bull. Torrey Bot. Club 5(3): 15. 1874. (Austin 1874; Staples & Imada 2006)
- =Radula excisiloba Steph., Bull. Herb. Boissier 5: 849. 1897. (Staples & Imada 2006)
Radula costaricensis Gottsche, J. Bot. 15: 226. 1877. (Polakowski 1877)
Radula crenulata Schiffn., Forschungsr. Gazelle 4(4): 21. 1889[1890]. (Schiffner 1889; Söderström et al. 2016)
†Radula cretacea Bechteler, M.A.M. Renner, Schäf.-Verw. & Heinrichs, Fossil Rec. 20(2): 206. 2017. (Bechteler et al. 2017)
Radula cubensis K. Yamada, J. Hattori Bot. Lab. 54: 241. 1983. (Yamada 1983)
Radula curvilobula M.L. So, J. Hattori Bot. Lab. 98: 176. 2005. (So 2005b)
Radula cuspidata Steph., Sp. Hepat. (Stephani) 4: 153. 1910. (Stephani 1912; Thouvenot et al. 2011)
- =Lejeunea dentata Mitt., Fl. Nov.-Zel. 2: 159. 1854. (Gibb et al. 2018) ≡Radula dentata (Mitt.) Mitt., Handb. N. Zeal. Fl. 2: 531. 1867, nom. illeg. ≡Stephanina dentata (Mitt.) Kuntze, Revis. Gen. Pl. 2: 839. 1891. ≡Radula dentifolia Grolle, Rev. Bryol. Lichénol. 32: 159. 1963. (Thouvenot et al. 2011)
Radula decora Gottsche ex Steph., Hedwigia 23: 145. 1884. (Stephani 1884a)
- =Radula dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26, Afd. 3(6): 61. 1900. (Hässel de Menéndez & Rubies 2009)
- =Radula inflata Steph., Sp. Hepat. (Stephani) 4: 211. 1910. (Hässel de Menéndez & Rubies 2009)
Radula decurrens Mitt., Fl. Vit.: 419. 1873. (Mitten 1873)
- =Radula recurviloba Steph., Sp. Hepat. (Stephani) 4: 186. 1910. (Grolle & Piippo 1984)
- =Radula micholitzii Steph., Sp. Hepat. (Stephani) 6: 512. 1924. (Grolle & Piippo 1984)
- =Radula multiflora var. reflexilobula Grolle & K. Yamada, J. Hattori Bot. Lab. 51: 326. 1982. (Yamada 1982a; Bardat et al. 2021)
Radula deflexilobula Promma, L.N. Zhang & R.L. Zhu, Cryptog., Bryol. 39(4): 484. 2018. (Promma et al. 2018)
Radula demissa M.A.M. Renner, PhytoKeys 27(Special issue): 53. 2013. (Renner et al. 2013c)
- =Radula epiphylla Colenso, Trans. & Proc. New Zealand Inst. 21: 71. 1889, nom. illeg. (Renner et al. 2013c)
Radula densifolia Castle, Rev. Bryol. Lichénol. 33: 385. 1965. (Castle 1965)
Radula diaphana K.I. Goebel, Organogr. Pfl. (ed. 2) 2(1): 677, 902. 1915. (Goebel 1915)
Radula diversifolia Steph., Sp. Hepat. (Stephani) 4: 212. 1910. (Stephani 1912)
- =Radula drepanophylla Steph., Sp. Hepat. (Stephani) 4: 212. 1910. (Hässel de Menéndez & Rubies 2009)
Radula dolabrata K. Yamada, J. Jap. Bot. 60: 1. 1985. (Yamada 1985a)
Radula eggersii K. Yamada, J. Hattori Bot. Lab. 82: 339. 1997. (Yamada 1997)
Radula elliottii Castle, J. Hattori Bot. Lab. 21: 12. 1959. (Castle 1959)
Radula emarginata K. Yamada & Piippo, Ann. Bot. Fenn. 26: 352. 1989. (Yamada & Piippo 1989)
Radula episcia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 318. 1885. (Spruce 1884–1885)
- =Radula cornucopiae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 318. 1885. (Spruce 1884–1885; Gradstein & Uribe-M. 2016)
- =Radula antilleana Castle, J. Hattori Bot. Lab. 21: 48. 1959. (Gradstein 2021)
Radula evansii Castle, Ann. Bryol. 11: 37. 1938. (Castle 1938)
Radula evelynae K. Yamada, J. Jap. Bot. 50: 115. 1975. (Yamada 1975a)
Radula falcata Steph., Hedwigia 23: 115. 1884. (Stephani 1884b)
Radula faurieana Steph., Sp. Hepat. (Stephani) 4: 207. 1910. (Stephani 1912)
Radula fendleri Gottsche ex Steph., Hedwigia 23: 146. 1884. (Stephani 1884a)
- =Radula dominicensis Steph., Sp. Hepat. (Stephani) 6: 507. 1924. (Schäfer-Verwimp 2010)
Radula fendleri Gottsche ex Steph. var. fendleri
Radula fendleri var. paroica F.R. Oliveira-da-Silva, Ilk.-Borg. & Gradst., Phytotaxa 454(1): 29. 2020. (Oliveira-da-Silva et al. 2020; Oliveira-da-Silva et al. 2021)
Radula fernandezana Steph., Kongl. Svenska Vetensk. Acad. Handl., n.s. 46(9): 84. 1911. (Stephani 1911)
Radula fissifolia Steph., Sp. Hepat. (Stephani) 6: 507. 1924. (Stephani 1925)
- =Radula tridens Steph., Sp. Hepat. (Stephani) 6: 516. 1924. (Thouvenot et al. 2011)
Radula flaccida Lindenb. & Gottsche, Syn. Hepat.: 726. 1847. (Gottsche et al. 1844)

≡Stephanina flaccida (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula flaccida var. propagulifera Gottsche, Mexik. Leverm.: 148 (244). 1863. (Gottsche 1863)
- =Radula epiphylla Mitt. ex Steph., Hedwigia 23: 151. 1884. (Wigginton 2018)
- =Radula foliicola Steph., Sp. Hepat. (Stephani) 4: 194. 1910. (Wigginton 2018)
Radula flaccida var. brachycalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 321. 1885. (Spruce 1884–1885; Söderström et al. 2016)
Radula flaccida Lindenb. & Gottsche var. flaccida
Radula flavifolia (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 259. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia flavifolia Hook. f. & Taylor, London J. Bot. 3: 476. 1844. (Hooker & Taylor 1844b)

≡Stephanina flavifolia (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula intempestiva Schiffn., Forschungsr. Gazelle 4(4): 20. 1889[1890]. (Hässel de Menéndez & Rubies 2009)
- =Radula cunninghamii Steph., Sp. Hepat. (Stephani) 4: 214. 1910. (Hässel de Menéndez & Rubies 2009)
Radula floridana Castle, Rev. Bryol. Lichénol. 36(1-2): 1. 1968[1969]. (Castle 1968b; Stotler & Crandall-Stotler 2017)
Radula forficata M.A.M. Renner, Austral. Syst. Bot. 26(4): 307. 2013. (Renner et al. 2013a)
Radula formosa (C.F.W. Meissn. ex Spreng.) Nees, Syn. Hepat.: 258. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia formosa C.F.W. Meissn. ex Spreng., Syst. Veg. [Sprengel] 4(2): 325. 1827. (Sprengel 1827b)

≡Stephanina formosa (C.F.W. Meissn. ex Spreng.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula spicata Mitt., Bonplandia (Hannover) 10: 19. 1862. (Söderström et al. 2011)
- =Radula pycnolejeunioides Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60(2): 247. 1893. (Grolle & Piippo 1984) ≡Radula formosa var. pycnolejeunioides (Schiffn.) N. Kitag., Acta Phytotax. Geobot. 30(1–3): 38. 1979. (Kitagawa 1979)
- =Radula caledonica Steph., Rev. Bryol. 35: 32. 1908. (Thouvenot et al. 2011)
- =Radula caledonica Steph., Sp. Hepat. (Stephani) 4: 232. 1910, nom. illeg. (Thouvenot et al. 2011)
- =Radula novae-guineae Steph., Sp. Hepat. (Stephani) 4: 233. 1910. (Thouvenot et al. 2011)
Radula fujitae Furuki, Bryol. Res. 9: 143. 2007. (Furuki 2007)
Radula fulvifolia (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 261. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia fulvifolia Hook. f. & Taylor, London J. Bot. 4: 85. 1845. (Hooker & Taylor 1845)

≡Stephanina fulvifolia (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula meyeri Steph., Hedwigia 27: 62. 1888. (Wigginton 2018)
- =Radula galapagona Steph., Sp. Hepat. (Stephani) 4: 176. 1910. (Stephani 1912; Oliveira-da-Silva et al. 2024)
- =Radula foreauana Beauverd, Sp. Hepat. (Stephani) 6: 514. 1924. (Yamada 1979a)
- =Radula schaefer-verwimpii K. Yamada, J. Jap. Bot. 65: 3. 1990. (Yamada 1990; Oliveira-da-Silva et al. 2024)
Radula gedena Gottsche ex Steph., Hedwigia 23: 146. 1884. (Stephani 1884a)
- =Radula gemmulosa S. Hatt., J. Hattori Bot. Lab. 4: 65. 1959. (Yamada & Iwatsuki 2006)
Radula gradsteinii K. Yamada, Trop. Bryol. 1: 37. 1989. (Gradstein & Florschütz-de Waard 1989)
Radula grandifolia Steph., Sp. Hepat. (Stephani) 4: 184. 1910. (Stephani 1912)
- =Radula douleana Steph., Sp. Hepat. (Stephani) 4: 184. 1910. (Castle 1968a)
Radula grandilobula Promma & Chantanaorr., Cryptog., Bryol. 36(3): 219. 2015. (Promma & Chantanaorrapint 2015)
Radula grandis Steph., J. Linn. Soc., Bot. 29(201): 271. 1892. (Stephani 1892)
Radula grevilleana Taylor, Ann. Mag. Nat. Hist. 20: 380. 1847. (Taylor 1847)
Radula grollei K. Yamada & Piippo, Ann. Bot. Fenn. 26: 379. 1989. (Yamada & Piippo 1989)
Radula guyanensis K. Yamada, Trop. Bryol. 1: 38. 1989. (Gradstein & Florschütz-de Waard 1989)
Radula hainanensis L.N. Zhang & R.L. Zhu, Bryologist 119(1): 53. 2016. (Zhang & Zhu 2016)
Radula hastata Steph., Sp. Hepat. (Stephani) 4: 163. 1910. (Stephani 1912)
Radula hattorii K. Yamada, J. Jap. Bot. 60: 259. 1985. (Yamada 1985a)
Radula hawaiica M.L. So, J. Hattori Bot. Lab. 98: 177. 2005. (So 2005b)
Radula hedingeri K.I. Goebel, Ann. Jard. Bot. Buitenzorg 7(1): 51. 1887. (Goebel 1887)
†Radula heinrichsii K. Feldberg, Schäf.-Verw., M.A.M. Renner, von Konrat & A.R. Schmidt, Cretaceous Res. 128(104987): 15. 2021. (Feldberg et al. 2021)
Radula helix (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 260. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia helix Hook. f. & Taylor, London J. Bot. 3: 475. 1844. (Hooker & Taylor 1844b)

≡Stephanina helix (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula magellanica Schiffn., Forschungsr. Gazelle 4(4): 21. 1889[1890]. (Hässel de Menéndez & Rubies 2009) ≡Stephanina magellanica (Schiffn.) Schiffn., Nat. Pflanzenfam. 1(3): 114. 1893.
- =Radula vagans Steph., Kongl. Svenska Vetensk. Acad. Handl., n.s. 46(9): 85. 1911. (Yamada 1993a)
- =Radula sainsburyana E.A. Hodgs. & Allison, Trans. & Proc. Roy. Soc. New Zealand 74: 279. 1944. (Renner et al. 2014)
- =Cololejeunea olivaceoviridis S.W. Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 7. 1958. (Arnell 1958; Váňa & Engel 2013)
Radula hicksiae K. Yamada, Cryptog. Bryol. Lichénol. 5: 191. 1984. (Yamada 1984a)
Radula holtii Spruce, J. Bot. 25: 209. 1887. (Spruce 1887)
Radula husnotii Castle, J. Hattori Bot. Lab. 21: 45. 1959. (Castle 1959)
Radula ilkiuborgesiae F.R. Oliveira-da-Silva & Gradst., Phytotaxa 653(1): 92. 2024. (Oliveira-da-Silva & Gradstein 2024)
Radula imposita M.A.M. Renner, PhytoKeys 27(Special issue): 65. 2013. (Renner et al. 2013c)
Radula inflexa Gottsche ex Steph., Hedwigia 23: 148. 1884. (Stephani 1884a)
Radula inouei K. Yamada, J. Hattori Bot. Lab. 45: 262. 1979. (Yamada 1979a)
†Radula intecta M.A.M. Renner, Schäf.-Verw. & Heinrichs, Earth & Environ. Sci. Trans. Roy. Soc. Edinburgh 107(2–3): 324. 2017. (Kaasalainen et al. 2017)
Radula involvens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 325. 1885. (Spruce 1884–1885)
Radula iwatsukiana K. Yamada, J. Hattori Bot. Lab. 58: 114. 1985. (Yamada 1985b)
Radula iwatsukii K. Yamada, J. Hattori Bot. Lab. 45: 275. 1979. (Yamada 1979a)
- =Radula scariosa var. nodulosa K. Yamada, J. Hattori Bot. Lab. 58: 127. 1985. (Yamada 1985b; Thouvenot et al. 2011)
Radula jamaicensis Pearson, Ann. Bryol. 4: 103. 1931. (Pearson 1931)
Radula jamesonii Taylor, London J. Bot. 5: 375. 1846. (Taylor 1846)

≡Stephanina jamesonii (Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula sprucei Steph., Sp. Hepat. (Stephani) 6: 515. 1924. (Gradstein & Uribe-M. 2016)
Radula japonica Gottsche ex Steph., Hedwigia 23: 152. 1884. (Stephani 1884a)

≡Stephanina japonica (Gottsche ex Steph.) Yasuda, Shokub. Kak.: 717. 1911.
- =Radula sendaica Steph., Sp. Hepat. (Stephani) 6: 514. 1924. (Yamada & Iwatsuki 2006)
Radula javanica Gottsche, Syn. Hepat.: 257. 1845. (Gottsche et al. 1844)

≡Stephanina javanica (Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula cordiloba Taylor, London J. Bot. 4: 375. 1846. (Renner 2014) ≡Stephanina cordiloba (Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula macrostachya Lindenb. & Gottsche, Syn. Hepat.: 726. 1847. (Gottsche et al. 1844; Gradstein & Uribe-M. 2016) ≡Stephanina macrostachya (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula wallichiana Lehm., Nov. Stirp. Pug. 10: 9. 1857. (Yamada 1979a)
- =Radula colliculosa Mitt., J. Proc. Linn. Soc., Bot. 5: 107. 1861[1860]. (Thouvenot et al. 2011)
- =Radula caldana Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33(7): 81. 1876. (Ångström 1876; Gradstein & Uribe-M. 2016)
- =Radula gottscheana F. Muell., Frag. Suppl.: 62. 1880, nom. illeg. (Renner 2014)
- =Radula sandei Steph., Hedwigia 23: 130. 1884. (Söderström et al. 2010)
- =Radula ceramensis Steph., Hedwigia 23: 132. 1884. (Yamada 1979a)
- =Radula ovalifolia Steph., Hedwigia 23: 135. 1884. (Yamada 1979a)
- =Radula surinamensis Steph., Hedwigia 23: 136. 1884. (Gradstein & Uribe-M. 2016)
- =Radula tenerrima Steph., Hedwigia 23: 136. 1884. (Castle 1968a)
- =Radula gracilis Mitt. ex Steph., Hedwigia 23: 147. 1884. (Gradstein 2021)
- =Radula amazonica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 321. 1885. (Spruce 1884–1885; Gradstein & Ilkiu-Borges 2009)
- =Radula amazonica var. campanensis Dumort., Trans. & Proc. Bot. Soc. Edinburgh 15: 322. 1885. (Spruce 1884–1885; Oliveira-da-Silva et al. 2021)
- =Radula amazonica var. negrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 322. 1885. (Oliveira-da-Silva et al. 2021)
- =Radula eggersiana Steph., Hedwigia 27: 302. 1888. (Stephani 1888; Castle 1968a)
- =Radula elegans Steph., Sp. Hepat. (Stephani) 4: 172. 1910. (Yamada 2000)
- =Radula falcifolia Steph., Sp. Hepat. (Stephani) 4: 174. 1910. (Schäfer-Verwimp 2010)
- =Radula longifolia Steph., Sp. Hepat. (Stephani) 4: 181. 1910. (Schäfer-Verwimp 2010)
- =Radula convexa Steph., Biblioth. Bot. 87: 232. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula lejeuneiformis Steph., Sp. Hepat. (Stephani) 6: 511. 1924. (Staples & Imada 2006)
- =Radula nietneri Steph., Sp. Hepat. (Stephani) 6: 512. 1924. (Yamada 1979a)
- =Radula similis Steph., Sp. Hepat. (Stephani) 6: 515. 1924. (So 2006)
- =Radula variabilis S. Hatt., Bull. Tokyo Sci. Mus. 11: 86. 1944. (Zhu & So 2001)
- =Radula unduliflora Castle, Rev. Bryol. Lichénol. 33: 387. 1965. (So 2006)
- =Radula cordiloba subsp. erigens M.A.M. Renner & Braggins, J. Hattori Bot. Lab. 97: 45. 2005. (Renner 2005) ≡Radula erigens (M.A.M. Renner & Braggins) M.A.M. Renner, Phytotaxa 81(1): 12. 2013. (Renner et al. 2013b; Renner 2014)
Radula jonesii Bouman, Dirkse & K. Yamada, J. Bryol. 15: 161. 1988. (Bouman et al. 1988)
Radula jovetiana K. Yamada, Cryptog. Bryol. Lichénol. 5: 193. 1984. (Yamada 1984a)
Radula juddii K. Yamada, J. Jap. Bot. 63(7): 241. 1988. (Yamada 1988)
Radula kilgourii M.A.M. Renner, Austral. Syst. Bot. 26(4): 313. 2013. (Renner et al. 2013a)
Radula kinabaluensis K. Yamada, Misc. Bryol. Lichenol. 6: 97. 1973. (Yamada 1973b)
Radula kitagawae K. Yamada, J. Hattori Bot. Lab. 58: 116. 1985. (Yamada 1985b)
Radula kojana Steph., Bull. Herb. Boissier 5: 105. 1897. (Stephani 1897a)

≡Stephanina kojana (Steph.) Yasuda, Shokub. Kak.: 718. 1911.
- =Radula decliviloba Steph., Sp. Hepat. (Stephani) 4: 153. 1910. (Yamada & Iwatsuki 2006)
Radula koponenii K. Yamada & Piippo, Ann. Bot. Fenn. 26: 364. 1989. (Yamada & Piippo 1989)
Radula kurzii Steph., Hedwigia 23: 153. 1884. (Stephani 1884a; Yamada 1979a)
- =Radula speciosa Gottsche ex Steph., Hedwigia 23: 155. 1884. (Stephani 1884a; Long & Rubasinghe 2014)
- =Radula andreana Steph., Sp. Hepat. (Stephani) 4: 182. 1910. (Yamada 1979a)
Radula lacerata Steph., Sp. Hepat. (Stephani) 4: 155. 1910. (Stephani 1912)
- =Radula farmeri Pearson, J. Linn. Soc., Bot. 46: 29. 1922. (Castle 1968a)
- =Radula laciniata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7: 193. 1931. (Herzog 1931; Castle 1968a)
- =Radula lacerata var. minor Hürl., Bauhinia 6: 301. 1978. (Hürlimann 1978; Thouvenot et al. 2011)
Radula laxiramea Steph., Sp. Hepat. (Stephani) 4: 178. 1910. (Stephani 1912)
Radula leiboldii Steph., Hedwigia 23: 116. 1884. (Stephani 1884b)
Radula lewisii K. Yamada, J. Hattori Bot. Lab. 74: 39. 1993. (Yamada 1993b)
Radula ligula Steph., Sp. Hepat. (Stephani) 4: 228. 1910. (Stephani 1912)
Radula lindenbergiana Gottsche ex C. Hartm., Handb. Skand. Fl. (ed. 9) 2: 98. 1864. (Hartman 1864; Hodgetts et al. 2020)

≡Radula complanata subsp. lindenbergiana (Gottsche ex C. Hartm.) R.M. Schust., Hepat. Anthocerotae N. Amer. 4: 609. 1980. (Schuster 1980)
- =Radula complanata f. plumulosa Nees, Naturgesch. Eur. Leberm. 3: 148. 1838. (Bernet 1888) ≡Radula complanata var. plumulosa (Nees) Debat, Fl. Musc.: 228. 1874.
- =Radula complanata f. propagulifera Nees, Naturgesch. Eur. Leberm. 3: 148. 1838. (Ros et al. 2007) ≡Radula complanata var. propagulifera (Nees) Debat, Fl. Musc.: 228. 1874.
- =Radula complanata f. tenuis Nees, Naturgesch. Eur. Leberm. 3: 148. 1838. (Bernet 1888) ≡Radula complanata var. tenuis (Nees) Debat, Fl. Musc.: 228. 1874.
- =Radula commutata Gottsche ex Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 58: 186. 1880. (Ros et al. 2007) ≡Radula lindenbergiana var. commutata (Gottsche ex Limpr.) C. Massal., Ann. Bot. (Rome) 1: 297. 1904.
- =Radula germana J.B. Jack, Flora 64(23): 355. 1881. (Ros et al. 2007) ≡Radula lindenbergiana var. germana (J.B. Jack) C. Massal., Ann. Bot. (Rome) 1: 297. 1904. ≡Radula lindenbergiana f. germana (J.B. Jack) Macvicar, Trans. & Proc. Bot. Soc. Edinburgh 25: 292. 1910.
- =Radula lindbergii Gottsche ex Lindb., Helsingfors Dagblad 1881(96): 3. 1881, nom. illeg. (Isoviita 1980)
- =Radula capensis Steph., Hedwigia 23: 131. 1884. (Wigginton 2018)
- =Radula ovata J.B. Jack ex Steph., Hedwigia 23: 154. 1884. (Ros et al. 2007)
- =Radula bornmuelleri Steph., Hedwigia 41: 277. 1902. (Ros et al. 2007)
- =Radula lindenbergiana f. propagulifera Schiffn., Lotos 60: 75. 1912. (Schiffner 1912)
- =Radula capensis f. gemmifera Sim, Trans. Roy. Soc. South Africa 15(1): 72. 1926. (Sim 1926)
Radula lindenbergiana var. atypa C. Massal., Mem. Accad. Verona 73(2): 23. 1897. (Massalongo 1897; Piippo 1990)
Radula lindenbergiana Gottsche ex C. Hartm. var. lindenbergiana
Radula lingulata Gottsche, Syn. Hepat.: 260. 1845. (Gottsche et al. 1844)

≡Stephanina lingulata (Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
Radula longicarinata F.R. Oliveira-da-Silva & Gradst., Phytotaxa 653(1): 94. 2024. (Oliveira-da-Silva & Gradstein 2024)
Radula longiloba K. Yamada, J. Hattori Bot. Lab. 54: 243. 1983. (Yamada 1983)
Radula longispica Steph., Sp. Hepat. (Stephani) 4: 183. 1910. (Stephani 1912)
Radula loriana Castle, J. Hattori Bot. Lab. 21: 6. 1959. (Castle 1959; Söderström et al. 2016)
Radula madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349. 1882. (Gottsche 1882)
- =Radula caespitosa Steph., Hedwigia 27: 107. 1888. (Renner et al. 2013a)
- =Radula exigua Steph., Sp. Hepat. (Stephani) 4: 195. 1910. (Grolle 2001)
Radula magna F.R. Oliveira-da-Silva & Gradst., Phytotaxa 653(1): 96. 2024. (Oliveira-da-Silva & Gradstein 2024)
Radula mammosa Spruce, Mem. Torrey Bot. Club 1: 127. 1890. (Spruce 1890)
- =Radula longituba Steph., Biblioth. Bot. 87: 233. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula verrucifolia Steph., Biblioth. Bot. 87: 233. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula lindigii Castle, J. Hattori Bot. Lab. 21: 29. 1959. (Gradstein & Uribe-M. 2016)
Radula marginata Gottsche, Lindenb. & Nees, Syn. Hepat.: 261. 1845. (Gottsche et al. 1844)

Replaced synonym: Jungermannia marginata Hook. f. & Taylor, London J. Bot. 3: 566. 1844, nom. illeg. (Hooker & Taylor 1844c)

≡Stephanina marginata (Gottsche, Lindenb. & Nees) Kuntze, Revis. Gen. Pl. 2: 839. 1891. ≡Radula tinctomarginata Grolle, Rev. Bryol. Lichénol. 29(1–2): 71. 1960, nom. illeg.
Radula marojezica E.W. Jones, J. Bryol. 17: 307. 1992. (Jones 1992)
Radula mauiensis M.L. So, J. Hattori Bot. Lab. 98: 178. 2005. (So 2005b)
Radula mexicana Lindenb. & Gottsche, Mexik. Leverm.: 150. 1863. (Gottsche 1863)
- =Radula ankefinensis Gottsche ex Steph., Hedwigia 23: 152. 1884. (Stephani 1884a; Gradstein et al. 2022)
- =Radula cordovana J.B. Jack ex Steph., Hedwigia 23: 163. 1884. (Gradstein & Costa 2003)
- =Radula claviflora Spruce, Mem. Torrey Bot. Club 1: 127. 1890. (Gradstein et al. 2003)
- =Radula holstiana Steph., Bot. Jahrb. Syst. 20: 320. 1895. (Grolle 2001)
- =Radula heteroica Steph., Sp. Hepat. (Stephani) 4: 167. 1910. (Wigginton 2018)
- =Radula autoica Steph., Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Kl. 88: 727. 1913. (Grolle 2001)
Radula microloba Gottsche, Syn. Hepat.: 259. 1845. (Gottsche et al. 1844)

≡Stephanina microloba (Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula microloba var. laxa Gottsche, Syn. Hepat. (fasc. 2): 259. 1845. (Gottsche et al. 1844; Hässel de Menéndez & Rubies 2009)
Radula microlobula Castle, J. Hattori Bot. Lab. 21: 35. 1959. (Castle 1959)
Radula minutilobula K. Yamada & Piippo, Ann. Bot. Fenn. 26: 377. 1989. (Yamada & Piippo 1989)
Radula mittenii Steph., Hedwigia 23: 148. 1884. (Stephani 1884a; Renner 2014)
Radula mizutanii K. Yamada, J. Jap. Bot. 48: 134. 1973. (Yamada 1973a)
Radula morobeana K. Yamada & Piippo, Ann. Bot. Fenn. 26: 358. 1989. (Yamada & Piippo 1989)
Radula multiamentula E.A. Hodgs., Rec. Domin. Mus. 4(11): 122. 1962. (Hodgson 1962; Söderström et al. 2016)
Radula multiflora Gottsche ex Schiffn., Forschungsr. Gazelle 4: 20. 1890. (Schiffner 1889; Bardat et al. 2021)
Radula myriopoda M.A.M. Renner, Austral. Syst. Bot. 26(4): 323. 2013. (Renner et al. 2013a)
- =Radula buccinifera var. fusiloba Steph., Hedwigia 28(4): 272. 1889. (Söderström et al. 2015b)
Radula neotropica Castle, J. Hattori Bot. Lab. 21: 31. 1959. (Castle 1959)
Radula nigra Pearson, J. Linn. Soc., Bot. 46: 31. 1922. (Pearson et al. 1922)
Radula nilgiriensis Udar & Dh. Kumar, J. Indian Bot. Soc. 61: 177. 1982. (Udar & Kumar 1982)
Radula norrisii K. Yamada & Piippo, Ann. Bot. Fenn. 26: 374. 1989. (Yamada & Piippo 1989)
Radula notabilis M.A.M. Renner, PhytoKeys 27(Special issue): 77. 2013. (Renner et al. 2013c)
Radula novae-hollandiae Hampe ex Lehm., Nov. Stirp. Pug. 7: 24. 1838. (Lehmann 1838)

≡Stephanina novae-hollandiae (Hampe ex Lehm.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
Radula novivrieseana K. Yamada, J. Hattori Bot. Lab. 51: 326. 1982. (Yamada 1982a)
Radula novocaledonica Hürl. & K. Yamada, J. Jap. Bot. 54: 238. 1979. (Hürlimann & Yamada 1979)
Radula novocaledoniensis K. Yamada, J. Hattori Bot. Lab. 58: 120. 1985. (Yamada 1985b)
Radula novoguineensis K. Yamada & Piippo, Ann. Bot. Fenn. 26: 360. 1989. (Yamada & Piippo 1989)
Radula nudicaulis Steph., Sp. Hepat. (Stephani) 4: 174. 1910. (Stephani 1912)
- =Radula goebelii Steph., Biblioth. Bot. 87: 232. 1916. (Herzog 1916; Gradstein & Uribe-M. 2016)
Radula nudicaulis var. delicatula P. Allorge & V. Allorge, Rev. Bryol. Lichénol. 19(1–2): 106. 1950. (Allorge & Allorge 1950; Hodgetts et al. 2020)
Radula nudicaulis Steph. var. nudicaulis
Radula nymanii Steph., Sp. Hepat. (Stephani) 4: 229. 1910. (Stephani 1912)
Radula obconica Sull., Manual: 688. 1848. (Sullivant 1848)
†Radula oblongifolia Casp., Schriften Königl. Phys.-Ökon. Ges. Königsberg 27: 4. 1887. (Caspary 1887)
- =†Radula bitterfeldensis Grolle, Feddes Repert. 96: 42. 1985. (Grolle & Meister 2004)
Radula obscura Mitt., J. Proc. Linn. Soc., Bot. 5: 107. 1861[1860]. (Mitten 1861b)
Radula obtusiloba Steph., Bull. Herb. Boissier 5(2): 105. 1897. (Stephani 1897a)

≡Stephanina obtusiloba (Steph.) Yasuda, Shokub. Kak.: 717. 1911.
- =Radula japonica var. major Amakawa, J. Hattori Bot. Lab. 13: 58. 1955. (Yamada 1986)
Radula obtusiloba Steph. subsp. obtusiloba
Radula obtusiloba subsp. polyclada (A. Evans) S. Hatt., J. Hattori Bot. Lab. 29: 275. 1966. (Hattori 1966; Stotler & Crandall-Stotler 2017)

Basionym: Radula polyclada A. Evans, Bull. Torrey Bot. Club 41(12): 607. 1914[1915]. (Evans 1914)
Radula oceania Castle, Rev. Bryol. Lichénol. 33: 390. 1965. (Castle 1965; Söderström et al. 2016)
Radula ocellata K. Yamada, J. Hattori Bot. Lab. 45: 209. 1979. (Yamada 1979a)
Radula okamurana Steph., Sp. Hepat. (Stephani) 4: 209. 1910. (Stephani 1912; Furuki & Mizutani 1994)
Radula onraedtii K. Yamada, Misc. Bryol. Lichenol. 8: 113. 1979. (Yamada 1979b)
Radula opaciuscula (Spruce) Castle, J. Hattori Bot. Lab. 21: 22. 1959. (Castle 1959)

Basionym: Radula episcia var. opaciuscula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319. 1885. (Spruce 1884–1885)
Radula oreopsis M.A.M. Renner, Telopea 17: 123. 2014. (Renner 2014)
Radula ornata E.A. Br. & Pócs, Telopea 9: 436. 2001. (Brown & Pócs 2001)
Radula ovalilobula K. Yamada, J. Hattori Bot. Lab. 45: 257. 1979. (Yamada 1979a)
Radula oyamensis Steph., Hedwigia 23: 149. 1884. (Stephani 1884a)

≡Stephanina oyamensis (Steph.) Yasuda, Shokub. Kak.: 717. 1911.
- =Radula oyamensis var. setulosa S. Hatt., J. Hattori Bot. Lab. 4: 69. 1950. (Yamada & Iwatsuki 2006)
Radula paganii Castle, J. Hattori Bot. Lab. 21: 33. 1959. (Castle 1959)
Radula pallens (Sw.) Nees ex Mont., Voy. Amér. Mér. 7(2): 71. 1839. (Montagne 1839)

Basionym: Jungermannia pallens Sw., Prodr.: 143. 1788. (Swartz 1788)

≡Stephanina pallens (Sw.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Jungermannia portoricensis Spreng., Syst. Veg. [Sprengel] 4(1): 226. 1827. (Sprengel 1827a; Castle 1968a)
- =Radula pallens var. minor Sw. ex Bisch., Fl. Azor.: 13. 1844. (Gottsche et al. 1844)
- =Radula pallens var. brasiliensis Gottsche, Lindenb. & Nees, Syn. Hepat. (fasc. 2): 256. 1845. (Gottsche et al. 1844)
- =Radula subsimplex Steph., Hedwigia 23: 130. 1884. (Gradstein & Lavocat Bernard 2020)
- =Radula kegelii Gottsche ex Steph., Hedwigia 23: 152. 1884. (Oliveira-da-Silva et al. 2021)
- =Radula didrichsenii Steph., Sp. Hepat. (Stephani) 4: 179. 1910. (Castle 1968a)
- =Radula obovata Castle, J. Hattori Bot. Lab. 21: 16. 1959. (Oliveira-da-Silva et al. 2021)
Radula pallens var. marginata F.R. Oliveira-da-Silva, Gradst. & Ilk.-Borg., Phytotaxa 564(1): 96. 2022. (Oliveira-da-Silva et al. 2022)
Radula pallens (Sw.) Nees ex Mont. var. pallens
Radula pandei Udar & Dh. Kumar, Lindbergia 9: 133. 1983. (Udar & Kumar 1983)
†Radula panduriformis M. Paulsen, D.J. Ohlsen, Cantrill & J.D. Stilwell, Rev. Palaeobot. Palynol. 338(105330): 3. 2025. (Paulsen et al. 2025)
Radula patens K. Yamada, Cryptog. Bryol. Lichénol. 5: 197. 1984. (Yamada 1984a)
†Radula patrickmuelleri K. Feldberg, Schäf.-Verw. & M.A.M. Renner, Fossil Rec. 25(1): 220. 2022. (Feldberg et al. 2022)
Radula peruviana K. Yamada, Beih. Nova Hedwigia 88: 79. 1987. (Schultze-Motel & Menzel 1987)
Radula philippinensis K. Yamada, J. Hattori Bot. Lab. 45: 299. 1979. (Yamada 1979a)
Radula physoloba Mont., Ann. Sci. Nat., Bot., sér. 2, 19: 255. 1843. (Montagne 1843)

≡Jungermannia physoloba (Mont.) Hook. f. & Taylor, Fl. Antarct. 1: 161. 1845. (Hooker 1844) ≡Stephanina physoloba (Mont.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula xanthochroma Colenso, Trans. & Proc. New Zealand Inst. 21: 70. 1888. (So 2005a)
- =Radula lycopodioides Colenso, Trans. & Proc. New Zealand Inst. 21: 71. 1889. (So 2005a)
- =Lejeunea ochracea Colenso, Trans. & Proc. New Zealand Inst. 21: 72. 1889. (Hamlin 1972)
- =Radula colensoi Steph., Sp. Hepat. (Stephani) 4: 215. 1910. (So 2005a)
Radula pinnata Pócs, Hattoria 11: 2. 2020. (Pócs 2020)
Radula pinnulata Mitt., Fl. Vit.: 410. 1871[1873]. (Mitten 1873; Yamada 1979a)
Radula plicata Mitt., Fl. Nov.-Zel. 2: 154. 1854. (Hooker 1855)

≡Stephanina plicata (Mitt.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
Radula pocsii K. Yamada, J. Hattori Bot. Lab. 54: 245. 1983. (Yamada 1983)
Radula portoricensis Steph., Hedwigia 27: 298. 1888. (Stephani 1888)
- =Radula guatemalensis Steph., Sp. Hepat. (Stephani) 4: 198. 1910. (Yamada 1986)
Radula prolifera Arnell, Ark. Bot. 13(2): 12. 1913. (Arnell 1913)
Radula protensa Lindenb., Bot. Zeitung (Berlin) 6: 462. 1848. (Meissner 1848)
Radula protensa var. erectilobula Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60(2): 247. 1893. (Schiffner 1893; Söderström et al. 2011)
Radula protensa Lindenb. var. protensa
Radula pseudoflaccida E.W. Jones, J. Bryol. 9: 501. 1977. (Jones 1977)
Radula pseudoscripta M.A.M. Renner, New Zealand J. Bot. 44: 340. 2006. (Renner 2006)
Radula pseudostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319. 1885. (Spruce 1884–1885)
Radula psychosis M.A.M. Renner, Austral. Syst. Bot. 26(4): 328. 2013. (Renner et al. 2013a)
Radula pugioniformis M.A.M. Renner, PhytoKeys 27(Special issue): 84. 2013. (Renner et al. 2013c)
Radula pulchella Mitt. ex Steph., Hedwigia 23: 149. 1884. (Stephani 1884a)
Radula punctata Steph., Hedwigia 23: 135. 1884. (Stephani 1884c; Oliveira-da-Silva et al. 2021)
- =Radula plumosa Mitt. ex Steph., Hedwigia 23: 154. 1884. (Stephani 1884a; Hässel de Menéndez & Rubies 2009)
- =Radula diversitexta Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26(III.6): 62. 1900. (Hässel de Menéndez & Rubies 2009)
- =Radula fruticosa Steph., Sp. Hepat. (Stephani) 4: 189. 1910. (Hässel de Menéndez & Rubies 2009)
- =Radula jackii Steph., Sp. Hepat. (Stephani) 4: 225. 1910. (Hässel de Menéndez & Rubies 2009)
- =Radula madothecoides Herzog, Rev. Bryol. Lichénol. 23: 55. 1954. (Hässel de Menéndez & Rubies 2009)
Radula pusilla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 320. 1885. (Spruce 1884–1885)
- =Radula mazarunensis K. Yamada, Trop. Bryol. 1: 38. 1989. (Gradstein & Florschütz-de Waard 1989; Gradstein 2021)
Radula quadrata Gottsche, Syn. Hepat.: 255. 1845. (Gottsche et al. 1844)

≡Stephanina quadrata (Gottsche) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula quadrata var. fusca Gottsche, Lindenb. & Nees, Syn. Hepat. (fasc. 5): 725. 1847. (Gottsche et al. 1844; Yamada 2000)
- =Radula affinis Lindenb. & Gottsche, Syn. Hepat.: 725. 1847. (Gottsche et al. 1844; Yamada 1993a) ≡Stephanina affinis (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula mollis Lindenb. & Gottsche, Syn. Hepat.: 725. 1847. (Gottsche et al. 1844; Stotler & Crandall-Stotler 2017) ≡Stephanina mollis (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula quadrata var. propagulifera Gottsche, Mexik. Leverm.: 144 (240). 1863. (Gottsche 1863)
- =Radula mollis var. prolifera Gottsche, Mexik. Leverm.: 145 (241). 1863. (Gottsche 1863)
- =Radula affinis var. propagulifera Gottsche, Mexik. Leverm.: 146 (242). 1863. (Gottsche 1863)
- =Radula quadrata var. manzanensis Gottsche, Ann. Sci. Nat., Bot., sér. 5, 1: 143. 1864. (Gradstein & Hekking 1979)
- =Radula andicola Steph., Hedwigia 23: 114. 1884. (Gradstein & Uribe-M. 2016)
- =Radula viridiaurea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 324. 1885. (Spruce 1884–1885; Yamada 1993a)
- =Radula recurvifolia Steph., Hedwigia 27: 63. 1888. (Wigginton 2018)
- =Radula calcarata Steph., Rev. Bryol. 36: 140. 1909. (Yamada 1993a)
- =Radula spongiosa Steph., Sp. Hepat. (Stephani) 4: 168. 1910. (Wigginton 2018)
- =Radula glauca Steph., Sp. Hepat. (Stephani) 4: 175. 1910. (Gradstein & Costa 2003)
- =Radula lescurii Austin ex Steph., Sp. Hepat. (Stephani) 4: 191. 1910. (Stotler & Crandall-Stotler 2017)
- =Radula pirottae Gola, Ann. Bot. (Rome) 13: 69. 1914. (Wigginton 2018)
Radula queenslandica K. Yamada, J. Hattori Bot. Lab. 62: 192. 1987. (Yamada 1987a)
Radula ratkowskyana K. Yamada, J. Jap. Bot. 59: 94. 1984. (Yamada 1984b; Renner 2005)
Radula recubans Taylor, London J. Bot. 5: 376. 1846. (Taylor 1846)

≡Stephanina recubans (Taylor) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
Radula reflexa Nees & Mont., Ann. Sci. Nat., Bot., sér. 2, 19: 255. 1843. (Montagne 1843)

≡Stephanina reflexa (Nees & Mont.) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula subsimilis Steph., Hedwigia 23: 136. 1884. (So 2006)
Radula renneri F.R. Oliveira-da-Silva, Ilk.-Borg. & Gradst., Nova Hedwigia 112(1–2): 128. 2021[2020]. (Oliveira-da-Silva et al. 2021)
Radula retroflexa Taylor, London J. Bot. 5: 378. 1846. (Taylor 1846; So 2005a)

≡Stephanina retroflexa (Taylor) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula miqueliana Taylor, London J. Bot. 5: 377. 1846. (Taylor 1846; Söderström et al. 2011) ≡Stephanina miqueliana (Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula fauciloba Steph., Sp. Hepat. (Stephani) 4: 188. 1910. (Thouvenot et al. 2011) ≡Radula retroflexa var. fauciloba (Steph.) K. Yamada, J. Hattori Bot. Lab. 45: 282. 1979. (Yamada 1979a)
- =Radula salakensis Steph., Sp. Hepat. (Stephani) 4: 205. 1910. (Söderström et al. 2010)
- =Radula helmsiana Steph., Sp. Hepat. (Stephani) 4: 231. 1910. (So 2005a)
- =Radula lunulatilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 226. 1934. (Horikawa 1934; Castle 1968a)
Radula rhombiloba Steph., Sp. Hepat. (Stephani) 4: 204. 1910. (Stephani 1912)
Radula robinsonii Steph., Sp. Hepat. (Stephani) 4: 214. 1910. (Stephani 1912)
- =Radula parvitexta Steph., Sp. Hepat. (Stephani) 6: 513. 1924. (Renner et al. 2013a)
Radula rupicola K. Yamada, J. Hattori Bot. Lab. 58: 124. 1985. (Yamada 1985b)
Radula saccatiloba Steph., Hedwigia 23: 129. 1884. (Stephani 1884c; Yamada 1987b)
Radula santacruziana K. Yamada & Gradst., Trop. Bryol. 4: 67. 1991. (Yamada & Gradstein 1991)
Radula scariosa Mitt., Bonplandia (Hannover) 9: 367. 1861. (Mitten 1861a)
- =Radula balansae Steph., Sp. Hepat. (Stephani) 4: 208. 1910. (So 2006)
Radula schofieldiana K. Yamada, J. Hattori Bot. Lab. 82: 337. 1997. (Yamada 1997)
Radula sharpii K. Yamada, J. Jap. Bot. 60: 4. 1985. (Yamada 1985a)
Radula sinskeana K. Yamada, J. Hattori Bot. Lab. 74: 41. 1993. (Yamada 1993b)
Radula sinuata Gottsche ex Steph., Sp. Hepat. (Stephani) 4: 161. 1910. (Stephani 1912)
- =Radula montana Steph., Sp. Hepat. (Stephani) 4: 176. 1910. (Gradstein & Uribe-M. 2016)
- =Radula obtusifolia Steph., Sp. Hepat. (Stephani) 4: 178. 1910. (Gradstein & Costa 2003)
Radula smithii F.R. Oliveira-da-Silva, Ilk.-Borg. & Gradst., Phytotaxa 564(1): 99. 2022. (Oliveira-da-Silva et al. 2022)
Radula socorana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475. 1947. (Gerola 1947)
Radula sonsonensis Steph., Sp. Hepat. (Stephani) 4: 201. 1910. (Stephani 1912)
†Radula sphaerocarpoides Grolle, Feddes Repert. 91: 404. 1980. (Grolle 1980; Heinrichs et al. 2016)

Replaced synonym: †Jungermannia sphaerocarpoides Casp., Schriften Königl. Phys.-Ökon. Ges. Königsberg 27: 1. 1887, nom. illeg. (Caspary 1887; Grolle & Meister 2004)
Radula splendida M.A.M. Renner & Devos, Nova Hedwigia 90(1–2): 113. 2010. (Renner et al. 2010)
Radula squarrosa K. Yamada, J. Jap. Bot. 65: 1. 1990. (Yamada 1990)
†Radula steerei Grolle, Mem. New York Bot. Gard. 45: 259. 1987. (Grolle 1987)
Radula stellatogemmipara C. Gao & Y.Huan Wu, Nova Hedwigia 80: 239. 2005. (Gao & Wu 2005)
Radula stenocalyx Mont., Ann. Sci. Nat., Bot., sér. 4, 3: 315. 1855. (Montagne 1855)
- =Radula pallens var. loborhiza Gottsche, Lindenb. & Nees, Syn. Hepat. (fasc. 2): 256. 1845. (Gradstein & Hekking 1989)
- =Radula tenella Gottsche, Mexik. Leverm.: 149. 1863. (Schäfer-Verwimp 2010)
- =Radula tubiflora Steph., Hedwigia 31: 172. 1892. (Wigginton 2018)
- =Radula vaginata Steph., Sp. Hepat. (Stephani) 4: 167. 1910. (Wigginton 2018)
Radula strangulata Hook. f. & Taylor, London J. Bot. 5: 377. 1846. (Taylor 1846; Renner et al. 2013c)

≡Stephanina strangulata (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
- =Radula levieri Steph., Sp. Hepat. (Stephani) 4: 227. 1910. (Renner et al. 2013c)
- =Radula silvosa E.A. Hodgs. & Allison, Trans. & Proc. Roy. Soc. New Zealand 74: 286. 1944. (Renner et al. 2013c)
- =Radula parviretis E.A. Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3: 87. 1965. (Renner et al. 2013c)
Radula striata Mitt. ex Steph., Hedwigia 23: 155. 1884. (Stephani 1884a)
Radula subacuminata L.N. Zhang & R.L. Zhu, Bryologist 124(2): 263. 2021. (Zhang et al. 2021)
Radula subinflata Lindenb. & Gottsche, Syn. Hepat.: 724. 1847. (Gottsche et al. 1844)

≡Stephanina subinflata (Lindenb. & Gottsche) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
Radula subsquarrosa S.W. Arnell, Ark. Bot., n.s. 4(1): 15. 1959. (Arnell & Mårtensson 1959)
Radula sullivantii Austin, Hepat. Bor.-Amer. Exsicc.: 22 (no. 88c). 1873. (Austin 1873)
Radula sumatrana Steph., Sp. Hepat. (Stephani) 4: 204. 1910. (Stephani 1912)
Radula tabularis Steph., Hedwigia 23: 131. 1884. (Stephani 1884c)
- =Radula indica Steph., Sp. Hepat. (Stephani) 6: 511. 1924. (Yamada 1979a)
- =Radula rara Steph., Sp. Hepat. (Stephani) 6: 514. 1924. (Castle 1968a)
†Radula tanaiensis K. Feldberg, Schäf.-Verw. & M.A.M. Renner, Fossil Rec. 25(1): 223. 2022. (Feldberg et al. 2022)
Radula tasmanica Steph., Sp. Hepat. (Stephani) 4: 212. 1910. (Stephani 1912)
Radula taylorii Steph., Hedwigia 23: 133. 1884. (Stephani 1884c)
Radula tectiloba Steph., Hedwigia 27: 298. 1888. (Stephani 1888)
- =Radula aurantii Spruce, Bull. Soc. Bot. France 36(Suppl. Congrés Bot. 1889): CXCIV. 1889[1890]. (Spruce 1889; Söderström et al. 2013)
- =Radula uleana Steph., Sp. Hepat. (Stephani) 4: 201. 1910. (Gradstein & Costa 2003)
- =Radula arsenei Steph., Sp. Hepat. (Stephani) 6: 505. 1924. (Gradstein & Costa 2003)
Radula tenera Mitt. ex Steph., Hedwigia 23: 149. 1884. (Stephani 1884a)
Radula tenuis K. Yamada, J. Hattori Bot. Lab. 54: 247. 1983. (Yamada 1983)
Radula thiersiae K. Yamada, J. Hattori Bot. Lab. 62: 198. 1987. (Yamada 1987a)
†Radula tikhomirovae Mamontov & Perkovsky, Ecol. Montenegrina 72: 192. . (Mamontov et al. 2024)
Radula tjibodensis K.I. Goebel, Ann. Jard. Bot. Buitenzorg 7(1): 53. 1887. (Goebel 1887)
- =Radula flavescens Steph., Sp. Hepat. (Stephani) 4: 203. 1910. (Grolle & Piippo 1984)
- =Radula reineckeana Steph., Sp. Hepat. (Stephani) 4: 225. 1910. (So 2006)
- =Radula tayabensis Steph., Sp. Hepat. (Stephani) 6: 516. 1924. (Castle 1968a)
Radula tokiensis Steph., Hedwigia 23: 150. 1884. (Stephani 1884a)

≡Stephanina tokiensis (Steph.) Yasuda, Shokub. Kak.: 718. 1911.
- =Radula kanemarui S. Hatt., J. Hattori Bot. Lab. 4: 67. 1950. (Wang et al. 2011)
Radula tonitrua Pócs & M.A.M. Renner, Telopea 24: 197. 2021. (Pócs & Renner 2021)
Radula underwoodii Castle, J. Hattori Bot. Lab. 21: 37. 1959. (Castle 1959)
Radula uvifera (Hook. f. & Taylor) Gottsche, Lindenb. & Nees, Syn. Hepat.: 258. 1845. (Gottsche et al. 1844)

Basionym: Jungermannia uvifera Hook. f. & Taylor, London J. Bot. 3: 392. 1844. (Hooker & Taylor 1844a)

≡Patarola uvifera (Hook. f. & Taylor) Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 410. 1877. ≡Stephanina uvifera (Hook. f. & Taylor) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula multicarinata Lindenb., Syn. Hepat. (fasc. 2): 258. 1845. (Gottsche et al. 1844; Castle 1968a) ≡Patarola multicarinata (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, Cl. Sci. Mat. 4: 410. 1877. ≡Stephanina multicarinata (Lindenb.) Kuntze, Revis. Gen. Pl. 2: 839. 1891.
Radula van-zantenii K. Yamada, J. Hattori Bot. Lab. 45: 260. 1979. (Yamada 1979a)
Radula varilobula Castle, J. Hattori Bot. Lab. 21: 19. 1959. (Castle 1959)
Radula venezuelensis K. Yamada, Misc. Bryol. Lichenol. 9: 122. 1982. (Yamada 1982b)
Radula ventricosa Steph., Sp. Hepat. (Stephani) 4: 187. 1910. (Stephani 1912)
- =Radula subpallens Steph., Sp. Hepat. (Stephani) 4: 203. 1910. (So 2006)
- =Radula coniloba Steph., Sp. Hepat. (Stephani) 6: 506. 1924. (Castle 1968a)
- =Radula ramicola Steph., Sp. Hepat. (Stephani) 6: 513. 1924. (Grolle & Piippo 1984)
Radula verrucosa K. Yamada, J. Hattori Bot. Lab. 45: 277. 1979. (Yamada 1979a)
Radula vieillardii Gottsche ex Steph., Hedwigia 23: 150. 1884. (Stephani 1884a)
Radula visianica C. Massal., Ann. Bot. (Rome) 1(4): 298. 1904. (Massalongo 1904)
Radula voluta Taylor ex Gottsche, Lindenb. & Nees, Syn. Hepat.: 255. 1845. (Gottsche et al. 1844)

≡Martinellius volutus (Taylor ex Gottsche, Lindenb. & Nees) Carruth., J. Bot. 3: 301. 1865. ≡Stephanina voluta (Taylor ex Gottsche, Lindenb. & Nees) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula ramulina Taylor, London J. Bot. 5: 374. 1846. (Taylor 1846; Gradstein & Uribe-M. 2016) ≡Stephanina ramulina (Taylor) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula macroloba Steph., Bull. Soc. Roy. Bot. Belgique 31: 121. 1892. (Grolle 2001)
- =Radula delessertii Steph., Bull. Soc. Roy. Bot. Belgique 32(2): 38. 1893. (Grolle 2001)
- =Radula lespagnolii Steph., Sp. Hepat. (Stephani) 4: 158. 1910. (Grolle 2001)
- =Radula newtonii Steph., Sp. Hepat. (Stephani) 4: 159. 1910. (Grolle 2001)
- =Radula stipatiflora Steph., Sp. Hepat. (Stephani) 4: 159. 1910. (Grolle 2001)
- =Radula subtropica Steph., Sp. Hepat. (Stephani) 4: 162. 1910. (Gradstein et al. 2003)
- =Radula allamanoi Gola, Mem. Reale Accad. Sci. Torino, ser. 2, 65: 8. 1914. (Gola 1914; Wigginton 2018)
- =Radula appendiculata Steph., Biblioth. Bot. 87: 231. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula boliviana Steph., Biblioth. Bot. 87: 231. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula grandiloba Steph., Biblioth. Bot. 87: 232. 1916. (Herzog 1916; Gradstein et al. 2003)
- =Radula multiramea Gola, Nuovo Giorn. Bot. Ital., n.s. 27: 246. 1920. (Wigginton 2018)
- =Radula dicksonii Castle, Rev. Bryol. Lichénol. 33: 343. 1965. (Yamada 2000)
- =Radula ramulina var. microphylla Jans, Proc. Kon. Ned. Akad. Wetensch., C 82: 428. 1979. (Jans 1979; Yamada 2000)
Radula vrieseana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 305. 1864. (Sande Lacoste 1864)
Radula weymouthiana Steph., Sp. Hepat. (Stephani) 4: 190. 1910. (Stephani 1912; Renner et al. 2013c)
Radula wichurae Steph., Sp. Hepat. (Stephani) 4: 168. 1910. (Stephani 1912; Hodgetts et al. 2020)
- =Radula limbata Schiffn., Oesterr. Bot. Z. 63: 442. 1913. (Ros et al. 2007)
Radula wrightii Castle, J. Hattori Bot. Lab. 21: 15. 1959. (Castle 1959)
Radula xalapensis Nees & Mont., Ann. Sci. Nat., Bot., sér. 2, 5: 56. 1836. (Nees von Esenbeck & Montagne 1836)

≡Stephanina xalapensis (Nees & Mont.) Kuntze, Revis. Gen. Pl. 2: 840. 1891.
- =Radula riparia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 322. 1885. (Spruce 1884–1885; Menzel 1984)
- =Radula frondescens Steph., Sp. Hepat. (Stephani) 4: 181. 1910. (Gradstein & Uribe-M. 2016)
Radula yamadae F.R. Oliveira-da-Silva & Ilk.-Borg., Nova Hedwigia 110(3–4): 288. 2020. (Oliveira-da-Silva & Ilkiu-Borges 2020)
Radula yangii K. Yamada, J. Hattori Bot. Lab. 45: 279. 1979. (Yamada 1979a; Yamada 1999)

Replaced synonym: Radula pinnulata B.Y. Yang, Taiwania 7: 36. 1960, nom. illeg. (Yang 1960)
Radula yanoella R.M. Schust., Phytologia 56: 72. 1984. (Schuster 1984)

A Synopsis of Radula

References (214)